UPWARD TRANSLOCATION OF FOODS IN WOODY PLANTS IO3 
Table 2. Philadelphus pubescens. Paired shoots measured from attachment to old stem. 
Rings made on old wood just below attachment of young shoots. May and June, 1919 
I 
Not Ringed. Leaves 
Remaining 
2 
Ringed. Leaves Re- 
maining 
3 
Not Ringed. Leaves 
Removed 
4 
Ringed. Leaves Re- 
moved 
Length 
in Cm. 
May 30 
Length 
in Cm. 
June 4 
Gain in 
Cm. 
Length 
in Cm. 
May 30 
Length 
in Cm. 
June 4 
Gain in 
Cm. 
Length 
in Mm. 
May 30 
Length 
in Mm. 
June 4 
Gain in 
Mm. 
Length 
in Mm. 
May 30 
Length 
in Mm. 
June 4 
Gain in 
Mm. 
a 
h 
c 
d 
e 
/ 
g 
19.0 
18.5 
17.0 
17-5 
25-5 
13.0 
23.0 
23.0 
12.5 
18.5 
20.0 
20.0 
15-5 
20.5 
35-5 
33- 5 
30- 5 
31- 5 
40-5 
30.0 
40-5 
42.0 
28.0 
35-0 
36.5 
38.5 
29.0 
34- 5 
17-5 
15-0 
13.5 
14.0 
17.0 
17-5 
19.0 
15.5 
16.5 
16.5 
18.5 
13.5 
14.0 
16.0 
14-5 
15.0 
18.5 
23.0 
20.5 
24.0 
24.0 
24-5 
21.5 
21.5 
24-5 
27-5 
27-5 
26.5 
27.5 
31.5 
35.0 
35-0 
38.0 
39-5 
40.0 
33-5 
35-0 
38.0 
41-5 
12.0 
12.5 
13.0 
8 0 
12.0 
14- 5 
14.0 
15.5 
15- 5 
12.0 
13.5 
13-5 
14.0 
4.0 
16.0 
13.0 
14-5 
19-5 
18.0 
17.0 
15.0 
16.0 
23.0 
16.5 
24-5 
24-5 
21-5 
10.5 
27.5 
20.5 
22.5 
22.5 
25.0 
-24.0 
25.0 
35.5 
19.0 
28.0 
30.0 
27-5 
6.5 
7.5 
8.0 
7-5 
4- 5 
8.0 
9.0 
9.0 
12.5 
2.5 
3-5 
5- 5 
6.0 
14.0 
12.5 
19.0 
16.5 
15.0 
19.0 
16.5 
20.0 
15-5 
15-5 
20.0 
20.5 
28.5 
^0.5 
15.5 
14.0 
21.5 
17.5 
20.5 
15-5 
21.5 
17.0 
17.0 
22.5 
22.5 
30.0 
27-5 
1-5 
1.5 
2.5 
I.O 
1-5 
2.0 
1-5 
1-5 
1.5 
2.5 
2.0 
10 
Ave. 
16.36 
12.96 
7.96 
1. 61 
Length 
in Mm. 
June 3 
Length 
in Mm. 
June 6 
Gain in 
Mm. 
Length 
in Mm. 
Junes 
Length 
in Mm. 
June 6 
Gain in 
Mm. 
Length 
in Mm. 
June 3 
Length 
in Mm. 
June 6 
Gain in 
Mm. 
Length 
in Mm. 
June 3 
Length 
in Mm, 
June 6 
Gain in 
Mm. 
h 
i 
k 
I 
32.0 
27.0 
28.0 
43-0 
42.5 
42.5 
42.5 
43-5 
39-5 
35-0 
36.0 
53.5 
54-0 
52.5 
52.5 
53-0 
7.5 
8.0 
8.0 
10.5 
11.5 
10. 0 
10. 0 
9-5 
38.0 
33-0 
41.5 
38.0 
42.5 
31-5 
40-5 
42.0 
45.0 
40.0 
49.0 
45.5 
51.0 
38.5 
47.0 
48.0 
7.0 
7.0 
7.5 
7-5 
8.5 
7.0 
6.5 
6.0 
45.0 
43-0 
36.5 
26.5 
44.5 
37-0 
46.0 
41.5 
47.5 
46.5 
40.0 
29.5 
48.0 
40.0 
50.5 
46.5 
2.5 
3-5 
3-5 
3-0 
3.5 
3-0 
4.0 
5.0 
24.0 
38.0 
36.0 
25.5 
35-0 
30.5 
28.5 
28.5 
25.5 
3.8.5 
37-5 
27.5 
36.5 
32.0 
30.0 
30.0 
1.5 
0.5 
1.5 
2.0 
1-5 
1.5 
1-5 
1-5 
Ave. 
9.38 
7.13 
1 3.56 
1.44 
The results of this second experiment are similar to those of the previous 
one. It is noticeable, however, that the ringed shoots with leaves (group 2) 
in this experiment show greater growth than shoots without leaves and 
not ringed (group 3), while in the previous experiment the reverse was true. 
In the experiment of 191 8 only a third to a half of the new shoot was above 
the ring or was leafless, while in that of 1919 the whole of each new shoot 
was above the ring. It seems very probable that the older leaves of the 
new shoot manufacture a large part of the food used in terminal growth 
after the shoot is well started. 
A similar experiment was tried with a tree of Northern Spy apple. 
This was started near the end of the growing season, which fact accounts 
for the small amount of growth obtained. All growth on this tree was 
practically completed by June 30, whereas with most of the other trees of 
the orchard it was nearly completed on June ii.^ 
2 The writer is much indebted to Prof. W. H. Chandler, of the department of pomology, 
who very kindly offered him the trees of the department orchard for experimentation, and 
to Prof. E. C. Auchter, now of the Maryland Agricultural Experiment Station, who assisted 
in these experiments with the apple. 
