UPWARD TRANSLOCATION OF FOODS IN WOODY PLANTS 
107 
data in table 5. In table 6 are given the freezing point lowerings of the 
terminal pair as well as of the first pair of stems below the terminal pair. 
These were determined for the same series but on a different day. 
Table 5. Philadelphus pubescens. Effect of ringing and removal of leaves on growth 
osmotic concentration, and soluble sugar content of the stem 
Length 
of 
Shoots 
in Cm. 
May 30 
Length 
of 
Shoots 
in Cm. 
June 4 
Gain 
in Cm. 
Length 
Ave. 
Gain 
per 
Pair. 
Freez- 
ing 
Point 
Depres- 
sion 
A 
Green 
Weight 
Total 
Dry 
Weight 
Mg. 
Invert 
Sugar 
per Gm. 
Green 
Weight 
Mg. 
Invert 
Sugar 
per Gm. 
Dry 
Weight 
I. Not ringed, leaves 
a 
19.0 
36.5 
33-5 
40.5 
42.0 
17.5 
15.0 
17.5 
19.0 
10. 0 
d 
18.5 
23.0 
23.0 
16.25 
18.75 
0.65° 
0.69° 
15.6 
1-74 
7.81 
II4.9 
2. Ringed, leaves pres- 
a 
16.0 
27-5 
26.5 
35-0 
38.0 
II.5 
12.0 
14.5 
14.0 
d 
14-5 
20.5 
24.0 
11-75 
14-25 
0.60° 
0.62° 
8.9 
12.0 
3. Not ringed, leaves 
removed 
a 
4.0 
16.0 
17.0 
15.0 
10.5 
27.5 
25.0 
24.0 
6.5 
8.0 
9.0 
d 
9.00 
8.50 
0.61° 
0.61° 
4-7 
4-9 
0.64 
4-5 
67.6 
4. Ringed, leaves re- 
moved 
a 
14.0 
12.5 
16.5 
20.0 
15-5 
14.0 
18.5 
21.5 
1.5 
1.5 
2.0 
1-5 
d 
1.50 
1-75 
0.455° 
0.52° 
2.7 
6.4 
0.445 
2.6 
52.2 
Table 6. Philadelphus pubescens. Effect of ringing and removal of leaves on osmotic con- 
centration of the stem as compared with the concentration of lower shoots of the same stem 
Total Gain for 
Each Pair 
of Shoots, 
May 30-June 4 
Green 
Weights 
A 
Green Weight 
of First Shoots 
Below 
A 
of First 
Shoots 
Below 
I. 
Not ringed, j 
((e) 
320 
9.5 
0.61 
10.2 
0.615 
leaves present 
(/) 
350 
12.93 
0.62 
4.4 
0.62 
2. 
Ringed, 
(ie) 
310 
16.8 
0.63 
8.23 
0.675 
leaves present 
Af) 
255 
8.8 
0.63 
12.9 
3- 
Not ringed. 
lie) 
215 
lO.O 
0.585 
I0.I3 
0.59 
leaves removed 
(/) 
60 
5.3 
0.61 
6.83 
0.57 
4- 
Ringed, 
(ie) 
30 
4-8 
0.53 
.5-3 
0.655 
leaves removed 
(/).. 
45 
6.3 
0.59 
0.66 
The cryoscopic data thus far obtained are not sufficiently extensive to 
warrant a detailed discussion, but they clearly show that the ringing of a 
defoliated shoot results in a distinct falling off of the osmotic concentration 
of that shoot. Whether the check in growth is due primarily to a lack of 
food necessary for energy or for building material, or to lack of water result- 
3 These are single shoots, not pairs. 
