200 
JOSEPH F. ROCK 
broadly sessile, partly stem-clasping, 5 to 12 cm. long, 11 to 30 mm. broad, 
5- to 1 1 -nerved; nerves converging near the base with the median costa; 
spikes I to 5, more than twice the length of the leaves, peduncles shorter 
than or equaling the leaves; flowers as in the species; stamens not exserted. 
Oahu: Gaud, in herb. Museum Paris; Gaud., Voy. Bonite, in Gray 
Herb.; U. S. Explor. Exped. in Gray Herb.; J. Remy, no. 427 in Gray 
Herb, ex herb. Museum Paris; Seemann, no. 2263 in Gray Herb.; Mann & 
Brigham, no. 85 in Gray Herb.; Manoa Valley, Rock, 1915, College of 
Hawaii Herb. no. 16001. 
PI. princeps var. Queleniana differs from the typical species in the simple, 
erect stem which is woolly throughout, while that of the species is glabrous. 
The leaves are not petiolate, but are more or less broadly stem-clasping. 
The stamens are also included, while in all the other varieties they are ex- 
serted. Bennett's Plantago Queleniana in the Berlin Herbarium belongs 
to PI. pachyphylla. Heller's ''PI. Queleana," no. 2610, and marked PL 
princeps in the Gray Herbarium has nothing in common with this variety; 
in fact, it represents a very anomalous form. 
In Hillebrand's collection in the Gray Herbarium there is a specimen 
collected in the Kohala mountains from the north coast of Hawaii, which he 
refers to Plantago princeps var. laxifolia, but which comes close to var. 
Queleniana; however, owing to the glabrous and somewhat petiolate leaves 
it is here omitted and referred to var. laxifolia. 
PI. princeps var. Queleniana is decidedly a montane variety and re- 
stricted to the rain forests, while the other varieties including the species 
occur on the outskirts of the forest and in the drier localities in the lower 
valleys. 
Plantago princeps var. elata Wawra Flora 32: 563. 1874. 
Shrub 2 m. high; stem undivided, erect, terete, foliose at the apex, 
otherwise naked, decidedly glabrous; leaves glabrous, lanceolate, 12.5 cm. 
long, 2.5 cm. broad, acuminate, narrowing at the base, and about 8 mm. 
broad at the insertion on the stem, glabrous, shiny above, 9-nerved, the 
2 or 4 inner nerves confluent below with the median nerve; spikes many 
(10 to 20), axillary, twice as long as the leaves; peduncle 7.5 to 10 cm. long, 
densely flowered, glabrous; bracts half the length of the calyx-lobes; 
flowers glabrous at the base; calyx lobes ovate-acute; tube of corolla equaling 
the calyx, the segments linear-lanceolate, acute, reflexed above the calyx; 
ovary ovate-obtuse; style filiform, long exserted; capsule oblong, twice as 
long as the calyx, long apiculate, circumscissile near the base, bilocular, 
locules one-seeded. 
Oahu: Mountains of Waianae, Wawra, no. 1728 h, in herb. Vienna; 
specimen not seen. 
This is not a synonym of Queleniana as cited by Drake del Castillo. 
It differs from Queleniana in the perfectly glabrous stem, which is even 
glabrous between the leaves. The stem is also unbranched. It is found 
in the dry regions of the Waianae mountains, while Queleniana is a montane 
species of the rain forest. 
