THE GENUS PLANTAGO IN HAWAII 
209 
Spikes remain. It differs, however, sufficiently from variety rotundifolia, 
especially in the thick lo-cm. long caudex and in the obovate-oblong leaves. 
Plantago pachyphylla var. glabrifolia Rock, Indig. Trees Haw. 
Isl. 77. 1913. 
Caudex very short, thick, and matted with wool; leaves forming large 
rosettes, broadly ovate in outline or ovate-oblong, acute, thick coriaceous, 
glabrous on both surfaces, broadly sessile at the base, 9- to ii-nerved, the 
nerves prominent below, the lateral ones arcuate, margins inconspicuously 
glandular-denticulate, 15 to 18 cm. long, 5.5 to 10 cm. broad; spikes i to 4, 
stout, glabrous or pubescent in the young stage, densely flowered in the 
upper third, flowers crowded toward the apex, loOsely flowered toward the 
base of the rachis ; bracts as long as t'he calyx, obtuse ; sepals acute ; corolla 
lobes acute or obtuse, reflexed; style long-exserted, gray-hairy; anthers 
more or less excised at the base, oblong to ovate; capsules unknown. 
Kauai: Waialeale, Oct. 20, 191 1, Rock, no. 8889 in College of Hawaii 
Herb. 
This variety differs from the others in the very broad, almost orbicular 
leaves which are glabrous on both surfaces, and in the glabrous spikes which 
are densely flowered, the flowers being oblong in outline rather than oval. 
With the exception of its glabrousness it would be referable to the typical 
Plantago pachyphylla var. mauiensis. 
Variety kauaiensis and its forma rohusta from Maui and Molokai 
undoubtedly are very closely related and are perhaps only forms of PL 
pachyphylla var. mauiensis, although large-leaved glabrous forms grow 
together with hirsute, slender and robust forms in the same locality. 
Plantago pachyphylla var. muscicola Rock. var. nova. 
Caudex short, thick, densely fibrillous below, the roots densely hairy as 
is the whole plant; leaves ovate to ovate-oblong, brittle, thick, fleshy, 
about 2 cm. or more in thickness, including the dense, gray hairs on both 
surfaces, these sometimes at right angles to the blade, 15 to 20 cm. long, 5 to 8 
cm. wide, acute at the apex, contracted below but broadly sessile at the 
base, 7- to 9-nerved ; nerves inconspicuous owing to the pubescence, arcuate ; 
margins of the leaves more or less conspicuously glandular-denticulate; 
spikes numerous (up to 10), densely hairy, stout, terete; bracts as long 
as the calyx or longer, hairy; sepals broadly ovate, acute, ciliate at the apex; 
corolla lobes broadly oval, acute, i-nerved ; capsule exserted, oblong, obtuse, 
2-seeded; seeds oval, dull brownish, rounded at both ends. 
Hawaii: Below summit of Kohala Mts. back of Waimea, elevation 
4,200 feet, in open bog, embedded in thick sphagnum, June, 1910, Rock, 
no. 8315 in the College of Hawaii Herb. 
This interesting variety differs from all the others in the thick, brittle, 
fleshy leaves which are densely hirsute on both surfaces with long, gray 
hairs which stand at right angles to the blade. The numerous spikes are 
very robust and hirsute as are the leaves. Otherwise as in Plantago pachy- 
phylla. Specimens of this variety have been distributed to herbaria as 
Plantago muscicola. 
