i86 
AMERICAN JOURNAL OF BOTANY 
[Vol. 8 
have no apparent perfume. At this point it may be well to mention that 
an aromatic perfume so similar as to be perhaps identical is noticed when 
the fresh end of a freshly broken branch is smelled, and that this perfume, 
unlike that of the flowers, is not confined to the male plant, but is observed 
also in the female. The similarity between the perfume of the sap and that of 
the flower becomes more marked upon purification. The "aqueous solution ' ' 
as made and described in a previous paper (i8) contains this more purified 
sap perfume. The panicles of the male and female flowers are somewhat 
differentiated as to location and structure. 
The flowering shoots of the male plant commonly bear as many flower 
panicles as leaves, in which case neither the highest leaves nor the lowest 
leaf develop any panicles in their axils. The lowest leaves of the flowering 
shoot soon fall off and more readily expose the flower panicles to insects, 
while the highest leaves remain and tend to protect the blossoms from the 
direct sunlight, wind, and rain. The panicles are 7 cm. long and stand 
somewhat stiffly upright at a sharp angle to the axil of the attached twig. 
The longer ones bear about a dozen side twigs of the first order, of which 
the three lowest ones are about 2 cm. long and in their turn are again richly 
branched. Toward the tip the side twigs of the first order become shorter 
and are not further branched. They are formed like a bunch of grapes, 
and the end of a panicle is likewise visibly terminated by a flower. The 
same regularity, as nearly as could be determined, appears in the arrange- 
ment of the side twigs of the first order on the panicle stem as was noticed 
in the phyllotaxy. Minute woolly hairs appear on the panicles at the 
blooming time, particularly on the bases of the panicle stem and on those 
of the side twigs. 
The flowers are placed singly on stalks from 4 to 7 mm. long, and have 
a diameter of from 5 to 7 mm. when fully opened. The flowers have 
5 calyx leaves, 5 petals, 5 stamens, and one rudimentary ovule; only by 
way of exception do 6 or 8 stamens occur, and in one flower with 6 stamens 
6 petals occurred also. 
The calyx leaves are tongue-shaped and have broad bases. They are 
about 2 mm. long and have a dark green color. 
The petals are long-elliptical in shape, narrowed at the base and at the 
point, and somewhat pointed in the front. They are 4 mm. long and in 
the middle about mm. wide. When in bloom the flowers are strongly 
bent downward. The color of the petals is light green, much lighter than 
that of the calyx leaves. 
The stamens are 2 3/^ mm. long. The white filaments, which are nearly 
twice as long as the anthers, shove themselves between the anther halves, 
which somewhat retreat from each other underneath. The anthers are 
introrse and are borne on upright but slightly curved filaments. 
The rudimentary ovary forms a keg-shaped pivot about i mm. high, 
and has 3 discernible stigmas. Between the ovary and the anthers is 
a disk, which during flowering time glistens with nectar. 
