June, 1921] CAMPBELL — THE EUSPORANGIATE FERNS 
of 0. Moluccanum Schlecht, which is possibly identical with 0. pedunculosum 
Desv. Specimens were secured in Ceylon of 0. reticulatum L. or some 
closely related species. These agreed closely with the species described 
by Mettenius, and indicated that in these tropical species of Euophio- 
glossum, the young sporophyte is absolutely destitute of any cauline tissues, 
being composed at first of a simple primary leaf, or cotyledon, which merges 
insensibly into the root (fig. i). A single central fibro-vascular bundle or 
"stele" extends without interruption from the petiole into the root, and its 
structure is essentially the same throughout, viz., "collateral" in the 
petiole, "monarch" in the root. 
Mettenius does not describe in detail the origin of the different organs 
of the embryo sporophyte. In 0. Moluccanum the writer found that the 
sporophyte at a very early stage consists of but two portions, a large basal 
Fig. I. Fig. 2. 
Fig. I. Median longitudinal section of a young sporophyte of OpMoglossum Moluc- 
canum, showing the primary stele traversing the cotyledon, L, and the root, R. Pr, the 
gametophyte. 
Fig. 2. Three longitudinal sections of the bud developed upon the primary root, r, 
of Ophioglosum Moluccanum. cot, cotyledon; the first two leaves of the bud; r\ the 
first root of the bud; st, the stem apex. 
foot and an apical conical portion developing subsequently into the coty- 
ledon. At this stage, the embryo is strongly suggestive of that of Anthoc- 
eros. The growing point of the primary root arises endogenously, being 
formed near the center of the embryo where the base of the young cotyledon 
