June, 1921] CAMPBELL THE EUSPORANGIATE FERNS 
As in Ophioglossum, the bundles are collateral, and, as is well known, 
there is developed a cambium between xylem and phloem, and also medul- 
lary rays, so that the structure of the woody cylinder in the older sporophyte 
is extraordinarily like that in the Gymnosperms and in many Dictoyledons. 
Fig. 3. A. Median longitudinal section of a young sporophyte of Botrychium Vir- 
ginianum, cut at right angles to the primary root. The two leaf-traces unite at the junction 
of the root. B. Transverse section of a similar sporophyte showing the two leaf-traces. 
C. Another section of the same sporophyte showing the junction of the two leaf-traces with 
the stele of the root. 
As in Ophioglossum, there is also in Botrychium no evidence of any 
procambial tissue in the axis above the youngest leaf-trace, i.e., the stelar 
tissues as in Ophioglossum are composed entirely of the coalescent leaf- 
traces. 
A still closer resemblance to Ophioglossum is shown by the young sporo- 
phyte of B. ohliquum Miihl., which the writer has recently examined. In 
this species the cotyledon and primary root, instead of forming an angle 
with each other, as in ^. Vir ginianum, have a common axis, and the orien- 
tation of these organs is like that in Ophioglossum Moluccanum. The 
writer is indebted to Dr. H. L. Lyon of Honolulu for the material upon 
which hfe investigations were made, and for the accompanying photograph 
(fig. 4)- 
Dr. Lyon^^ first directed attention to the peculiarities of this species, 
Lyon, H. L. A new genus of Ophioglossaceae. Bot. Gaz. 40: 455-458. 1905. 
