Oct., 1921] BECHTEL — FLORAL ANATOMY OF URTICALES 
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parenchymatous tissue. Each stele is composed of four strands. From 
these strands there pass off in the base of the flower four traces to the 
perianth (PI. XIX, figs. 11, 12, p)] the remaining four continue into the 
pistil (figs. II, 12, I, i\ 0). The perianth parts become distinct at a level 
with the ovule (fig. 16), which is not far above the level where the flowers 
become distinct from each other (fig. 13). 
The perianth parts vary much in the amount of vascular supply. In 
addition to the one main bundle, there frequently exist in the same inflores- 
cence flowers having perianth parts with few to many small, weak bundles 
(PI. XIX, figs. 14-17, n). The peculiar feature of these bundles is that they 
cannot be followed to their origin because of the lack of any organization 
suggesting bundles in the lower part of the perianth. These small strands 
are either branches of the main bundle of the perianth part, or, as shown in 
Ulmus, they are a separation of the strands that are now passing off from 
the floral axis as a common trace. The presence and abundance of these 
faint bundles vary according to the crowding of the flowers in the dense 
capitate inflorescence. The two lateral strands to the pistil (PI. XIX, fig. 
12, 0) approach each other, becoming one strand (figs. 13-16, 0 + 0) which 
passes to the posterior side upward. into the pendulous ovule (fig. 18, 0). 
The anterior bundle passes up the anterior side of the pistil to the tip of the 
single filiform style. This is the bundle to the anterior carpel (figs. 12-18, i). 
The corresponding bundle passes up three-fourths of the height of the ovary 
(figs. 12-18, i^, on its posterior side. Comparing the vascular supply of the 
two carpels of Morus with that of Madura, the conclusion is that the 
posterior carpel of Madura is abortive. This abortion of the carpel and 
the non-actinomorphic condition of the flower make zygomorphy a feature 
of the Madura flower. 
In the staminate flower four traces from the pedicel separate into 
strands to the perianth parts and to the stamens. The carpels are sup- 
pressed, and there are no signs of any vascular tissue in the central portion 
of the flower (PI. XIX, fig. 19). 
Cannabis sativa L. The pedicel of the pistillate flower has four stelar 
strands (PL XX, fig. 2). One of these strands (fig. 2, a) passes off anteriorly 
into the bract which completely envelops the flower (figs. 2-4, hr). The 
three remaining strands pass up into the pistil of the flower. Two strands 
(PI. XX, figs. 2, 3, I, i^) which have nearly the same size pass up the dorsal 
sides of the two carpels to the tips of the two styles respectively (figs. 2-8, 
I, l^). The fourth strand of the pedicel, which is opposite the strand passing 
into the enveloping bract, is twice the size of any of the other strands 
(PI. XX, figs. 2-4, 0). This strand maintains its bulkiness as it passes up 
posteriorly into the pendulous ovule (figs. 5-7, 0). 
A transverse section of the flower just below the ovule shows six dis- 
tinctly Hgnified bundles in the perianth (PI. XX, fig. 5, m). The appearance 
of the tissues of the perianth suggests more bundles than those having 
