Oct., I92I] EMERSON ABERRANT CHROMOSOME BEHAVIOR 
since the second sperm nucleus carried C and not R and the polar nuclei R 
and not C, if the two elements divided without undergoing fusion, C and R 
could never have been brought together and no aleurone color could have 
developed in any part of the seeds. 
Taei.e I. Aberrant seeds of maize from crosses of colorless waxy, c wx, hy colored corneous ^ 
C Wx " 
Pedigree No. 
of 9 Parent 
9062- 2. 
3 
3- 
6. 
9064- 2. 
9491- I. 
2. 
10168- 5. 
10169- 3. 
10170- I. 
2 . 
2. 
4. 
5- 
7- 
10171- I. 
4- 
6. 
7- 
8. 
10. 
II . 
II . 
13- 
14- 
16. 
17- 
18. 
18. 
19. 
23- 
24. 
25- 
27. 
29. 
10182- I. 
5- 
II . 
29 ears. . 
Approximate 
Number of 
Normal Seeds 
Number of Aberrant Seeds. Endosperm under 
Colorless Parts 
460 
600 
500 
280 
10 
560 
250 
480 
430 
400 
300 
460 
390 
310 
70 
460 
450 
650 
620 
340 
430 
290 
540 
160 
560 
530 
270 
630 
470 
190 
240 
350 
140 
470 
430 
150 
240 
120 
9,580 
Waxy 
Corneous 
Undetermined 
Total 67 ears . 
23,810 
55 
In a similar way the writer (Emerson, 191 5) was able to show that 
Webber's second hypothesis is incorrect. Two colorless types, each having 
only one of the two complementary aleurone-color factors C and R, were 
crossed as in the experiments of East. In addition, the type used as the 
female parent was sugary and the one used as the male parent was starchy. 
