108 
Iiihet'ltatice in the Foxglove 
Two of these plants of 5(f (G 2 and 7) were selfed, and the generation raised 
exhibited a lowered pelorism. The various selections made and the results 
obtained are shown in the accompanying table. It will be seen that finally on 
the selling of plant G 7, 10, 20, 4 (0 ') only non-peloric offspring were obtained. 
2. General Coloration of the Corolla. 
As described in the previous report {loc. cit.) the intensity of the purple 
coloration was measured by comparing it with a colour-scale founded on the 
intensity of colour by transmitted light of varying depths of a standard colour- 
solution. 
Purple and white foxgloves exhibit the ordinary Mendelian relationship, purple 
being dominant. A confusing aspect of the problem is introduced by the fact 
that " white " foxgloves are not necessarily entirely white, since they may exhibit 
a faint purple coloration which on the colour-scale adopted may amount to 
about 5. On crossing such a plant with an ordinary purple plant segregation 
occurs when the hetcrozygcjus offspring are self-fertilised. Any higher coloration, 
say 10 — 15, does not exhibit segregation, but gives a blended inheritance, and 
such a plant is to be regarded as a very pale purple one and not " white." From 
certain observati(jns that have been made it is probable that a similar condition 
occurs in the Blue Agapanthus lily, since some of the " white " plants have flowers 
faintly tinged with blue. It is quite likely that the phenomenon is general, and it 
may throw an important light on the physical theory of heredity. Possibly it 
may be surmised that a factor for a coloration of less than 5 units is unable 
to blend with, or influence, the factor controlling a higher coloration, in that we 
have reached the lowest dynamic unit. 
Of the ten original plants, five were purple and homozygous, four were; purple 
and heterozygous and one was white or recessive. These were very variously 
crossed in all manner of ways. In the accompanying table the experimental 
results are compared with the Mendelian expectation for the different gametic 
pairings. 
General Guloratioii of Corolla — Breeding ResulU. 
Gametic Nature 
Number 
Number 
White 
Purple 
of 
of 
of 
Pairings 
Families 
Offspriiij; 
Experimental 
Expectation 
Experimental 
Expectation 
DDy.DD 
120 
1620 
2 + 3 
0 
1615 
1620 
RR X RR 
17 
336 
330 
336 
6 
0 
DR X DR 
50 
785 
190 
196 
595 
589 
DR X DD 
11 
103 
0 
0 
103 
103 
DR X RR 
8 
76 
24 
38 
52 
38 
DD X RR 
8 
87 
0 
0 
87 
87 
Totalw 
214 
3007 
549 
570 
2458 
2437 
