126 
Inheritance in the Foxglove 
With reference to the characters which blend, the accompanying table sum- 
marizes the results obtained for parental coiTclation. Mid-parents and self- fertilised 
parents are regarded as comparable. 
Character 
Number of 
Offspring 
Coefficient of 
Correlation. 
Parents and 
Offspring 
Intensity of pelorism (homozygous recessive,) 
mid-parents and self-fertilised parents) j 
Intensity of general purple coloration (homo-\ 
zygous dominants, self-fertilised parents) J 
Seed-length (self-fertilised parents) 
Spotting (self-fertilised parents) 
Ratio of Corolla (self-fertilised parents) 
530 
529 
46 
716 
713 
•520 
•707 
•378 
■560 
•601 
The probable errors of these results are reasonably small and the average 
coefficient for the 5 characters is "553 which is not far removed from the average 
coefficient found by Professor Karl Pearson for a large number of characters in a 
variety of different organisms. 
It must be again emphasized that these results are based on self-fertilised 
generations of pedigree plants of known gametic constitution, and on Johannesen's 
theory of pure-lines these parental coefficients should be zero, or at least very 
small. 
The evidence of the present investigation is therefore definitely against any 
general application of the theory of pure-lines and of genotypes of any appreciable 
magnitude, and further it indicates that selective breeding within self-fertilised 
generations of a homogeneous race is capable of modifying that race to a marked 
degree. 
EXPLANATION OF PLATE I. 
Figs. 1 and 2. — Pelorism o£ maximum intensity ; grade 100°. Corollas absent, sessile anthers. 
Figs. 3 and 4. — Perfect pelorism, grade 100°. Corollas joined along their split edges forming a complete 
saucer. Stamens with filaments. 
Fig. 5. — Peloric flower of side-axis ; the axis terminates in an ovary. 
Fig. 6. — Pelorism of grade 100°. Numerous flowers fused irregularly forming a rosette, the axis has 
grown through the crown. 
Figs. 7 and 8. — Incomplete pelorism of main axes, grade 75°. A spiral bending often occurs. 
Fig. 9. — Faintly defined pelorism. When such occurred on the lateral axes the plant was said to 
possess a grade of 25°. Side view, and view from above. 
Fig. 10.— Flowering axis of a conspicuous sport in which practically all the corollas are completely split 
longitudinally into four elongated blades. Nature of inheritance obscure. 
The photographs were kindly taken by Dr Conrad Akerman. 
