Apparent Apogamy in Pterygodium Newdigatoe. 597 
It seems probable that there is some relationship between the extraordinary 
polymorphism of this species and its apparent apogamy. 
The exact nature of the nuclear changes in the female elements will be 
investigated after a complete series of the various stages has been obtained. 
The above investigation was commenced with the assumption that the 
gland-like bodies at the tips of the rostellum arms are reduced anther cells. 
An examination of the closely allied species, Pterygodium catholicum, has 
suggested the possibility that the masses of tissue in question may represent 
the rostellary glands. On dissecting away the anther from young flowers of 
Pt. catholicum, the appearance presented by the rostellary arms is strikingly 
like that of the two-armed form of Pt. Newdigatae. A microscopic examina- 
tion of the gland-like tissues in each case reveals a further similarity. If 
this supposition should prove to be correct, then we have in the cleistogamous 
form of Pt. Newdigatae complete suppression of the fertile anther together 
with the lip appendage, which is considered by some as representing the 
posterior stamen of the inner whorl. 
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