A Coutiibution to our Knoiuledge of the Anatomy of Cliamacleonti. 91 
resemblance to dilejpis rather than to the i:)iimilii,s section in the total 
absence of pigmentation on the peritoneum, which is black in damaranus, 
ventralis, etc. We have examined several specimens each of lymnilus, 
vmtralis, aad daniaranus in every case finding no trace of diverticula. 
As the piiviilus section is fairly homogeneous in external characters, we 
assume that the absence of diverticula is characteristic of the whole sec- 
tion. Further, Brookesia of Madagascar is in agreement with the pumilus 
section in this respect. It would be of great interest to know what are the 
lung characters of the genus Bhamijlioleon of tropical Africa, but unfortu- 
nately no material is available. We venture to surmise, however, that it 
will prove to be similar to Brookesia. Chamaeleon tigris of the Seychelles 
also seems to be allied to the i:)innilus section, but we do not know its 
lung characters. 
(Through the courtesy of Mr. G. A. Boulenger, F.R.S., we have been 
enabled to examine a specimen of Bhamplioleon spectrnvi (Oban, Calabar) 
and two specimens of Chamaeleon tigris (Silhouette Is., Seychelles) in the 
British Museum collection. 
The lungs of Bhampholeon were found to be simple, without trace of 
diverticula. Both the specimens of Chamaeleon tigris showed that the 
lungs were without diverticula such as those described below for other 
species of Chamaeleon ; but the lungs in both had ventrally and at a point 
about two-thirds the length of the organ distally a rounded lobe supplied 
with blood-vessels. In the absence of true diverticula C. tigris resembles 
the pumilus section of the Cape ; on the other hand, the examination of a 
rather badly preserved skull of this species inclines us to the view that it 
is in all probability an isolated form, related most nearly perhaps to some 
of the Madagascar species.) t 
Undoubtedly the primitive scheme of lung of the Rhiptoglossa is that 
which conforms to a great extent to that of other Lacertilia ; and for this 
reason in the genus Chamaeleon, the Cape species, C. pumilus, ventralis, 
damaranus, etc., seem to be (they may be degraded) nearer the primitive 
Rhiptogiossan stock than the Malagasy and other species of this genus 
(cf. Beddard, loc. cit. p. 41, line 44), and similarly the genus Brookesia 
must rank as the Malagasy representative of that primitive stock, though 
it is now in some respects a highly specialized creature. 
The variation which obtains in the peculiar type of lung as presented 
by the common Chamaeleon and by C. dilepis may be illustrated by refer- 
ence to the following species : — 
Chaniaeleoji dilepis, Leach, var. quilensis, Boc. (Text-fig. 1). — A good 
deal of variation obtains, and even in the same individual the number of 
* Our specimens of damaranua came from Knysna. It is highly improbable that the 
original locality record for this species, " Damaraland," can be correct, 
t Inserted April, 1914. 
