AMERICAN 
JOURNAL OF BOTANY 
Vol. Ill June, 1916 No. 6 
THE ARCHEGONIUM AND SPOROPHYTE OF TREUBIA 
INSIGNIS GOEBEL 
Douglas Houghton Campbell 
One of the largest and most interesting liverworts is Treubia 
insignis discovered by Goebel in western Java, and named by him 
for the late distinguished director of the famous botanical gardens at 
Buitenzorg in Java, Dr. Melchior Treub. 
The plant was collected near Tjibodas on Mt. Gedeh, a volcano in 
western Java, and it has since been found repeatedly by various 
botanists in this neighborhood. Schiffner^ gives this as the only 
known habitat, but later collectors have found the plant (or a closely 
related species) in several widely separated regions. Goebel himself 
collected a Treubia in New Zealand, and it has been reported from 
Tasmania, Tahiti, Samoa and Patagonia. 
''Stephani"2 recognizes two species, T. insignis, from Java and 
Tahiti, and T. hracteata from Samoa. Sterile material only has been 
found in this latter species. T. hracteata has recently been reported 
from Tasmania,^ and it is not unlikely that the New Zealand species is 
the same. 
In May, 191 3, the writer collected a single specimen of Treubia on 
Mt. Banajao in Luzon, Philippine Islands. The specimen was sterile, 
but except for its somewhat smaller size, it seemed to be identical with 
material collected in Java. 
During a visit to Java in 1906 the writer secured a large amount 
of material near Tjibodas, where the plant was found growing in some 
places in great profusion on the ground and on rotten logs. Only a 
few plants with sporogonia could be found, but a number of plants 
^ Die Hepaticae der Flora von Buitenzorg. Leiden. 1900. 
2 Stephani, F. Species Hepaticarum. Mem. Herb. Boiss., 16. 1900. 
' Rodway, L. Notes on Treubia insignis Goebel. Papers Proc. Roy. Soc. 
Tasmania. 
(The Journal for May (3: 211-260) was issued May 26, 19 16.] 
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