328 
GERTRUDE E. DOUGLAS 
CORTINARIUS ARMILLATUS 
(Figs. 19-24) 
Cortinarius armillatus and Cortinarius distans are of especial 
interest on account of the fact that both belong to the only sub- 
genus, Telamonia, described as possessing a universal veil. C. armil- 
latus derives its name from the fact that, as the veil breaks away from 
the margin of the pileus, it is left on the stem in a series of rings. 
The material was not found in great abundance but represents the 
most critical stages of development, except the origin of the hymeno- 
phore. The youngest stage (fig. 19) already shows a differentiation 
into three regions. In the center there is a region of active growth, 
conical in shape, which probably represents the stem fundament. 
This is surrounded by a zone of ground tissue, on the surface of which 
is a layer of large thick-walled hyphae. The latter is probably not a 
protoblem, evidence of which shows in the older fruit bodies, but is 
perhaps due to changes in the hyphae, caused by some substance in 
the substratum with which it came in contact. The older specimen, 
represented in figure 20, shows a considerable increase in the central 
growth zone, which now extends upwards nearly to the apex of the 
young basidiocarp. Progression of the growth area of the stem 
fundament upward gives rise to the pileus fundament which is sur- 
rounded laterally by loose ground tissue and blematogen, there being 
evidence of a slight but broad constriction between pileus and stem 
fundament. This method of differentiation of stem and pileus funda- 
ment is like that described by A. MoUer^^ (p. 70) for Rozites gongylo- 
phora, and by Atkinson^ ^ for Lepiota cristata and seminuda. 
On the outside is a very dehcate layer of fibrils, which in all prob- 
abihty here represents a true protoblem. It is present in all the older 
stages where we find very delicate fibrils or scales gradually being 
shed by the plant. For this reason, we may assume that it is also 
present in the early stages, but was lost from the button of figure 19 
during the preparation processes. The zone just within represents the 
blematogen, a region very indefinite in the young stages. As the plant 
becomes older, the boundaries of the blematogen become more distinct 
1^ Moller, A. Die Pilzgarten einiger siidamerikanischer Ameisen. Bot. Mit- 
theil. Trop. 6: 1-127, figs. 1-4. pis. 1-7. 1893. 
1^ Atkinson, Geo. F. The development of Lepiota cristata and L. seminuda. 
Proc. 20th Anniversary N. Y. Bot. Gard. 
