A STUDY OF DEVELOPMENT IN THE GENUS CORTINARIUS 329 
(figs. 21 and 22) until, in the mature stages, it becomes a very definite 
area. Its duplex character shows even more distinctly here than in 
C. anfractus. The outer layer is continuous completely over the 
surface of the pileus, the gill cavity and the upper part of the stem. 
It is considerably firmer than the floccose portion within, which 
extends from the upper surface of the pileus margin to the stem, out- 
side the partial veil. The latter is very fragile and as the epinastic 
growth of the pileus takes place, the hyphae are considerably stretched 
and become united in strands (fig. 23). The partial veil gradually 
breaks loose from the pileus margin and, together with the portion of 
the blematogen outside with Vv^hich it remains in contact, is torn 
apart by the elongating stem so that the series of delicate rings, above 
described, are left around the stem. The stages of early development 
of the gills are lacking but figures 21 and 22 show that they are pre- 
ceded by a palisade layer and extensive gill cavity. Figure 24 repre- 
sents a later stage of the gills and shows the origin of a forked gill, 
by means of a secondary salient growing out at the base of a primary 
gill. 
CORTINARIUS DISTANS 
(Figs. 63-69) 
A few photographs of Cortinarius distans, also belonging to the 
sub-genus Telamonia, are shown in figures 63-69. These were chosen 
to illustrate the character of the veil and the method of development 
of the gills. A very distinct blematogen layer covers the fruit body. 
Its hyphae are characteristically large and generally radial in their 
arrangement over the apex of the fruit body, but become nearly 
parallel at the sides. They take the stain very lightly. The blemato- 
gen soon breaks into scales and becomes easily removed, exposing 
the surface of the pileus, made up of very firm pseudoparenchymatous 
tissue (fig. 69). The pileus retains its conical shape as it pushes 
through the soil and on this account, the blematogen first disappears 
from the apex and much later at the margin of the pileus. Its floccose 
character is shown in figure 64, an enlargement of figure 63. On 
account of the scaling off of the outer layers, it was not determined 
whether or not the blematogen was originally duplex. 
The method of gill formation is like that of C. anfractus, with the 
exception that the salients are broader and more widely distant, 
resulting in a triangular gill at maturity. On account of this feature, 
the plant has received its name of C. distans. 
