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fact that, of these forms, only Puccinia Circaeae has been collected in 
America in sufficient numbers to render its status definite. There is 
a considerable variation, in the morphological characters, between 
these different short-cycled forms. These characters appear to 
indicate evident relationships or correlations with different long-cycled 
species of rust upon the same or similar hosts. As is indicated under 
each species, and discussed further on in this article, these short-cycled 
forms appear to fall into two, very doubtfully three, general groups: 
the first, represented by Puccinia Circaeae, Puccinia gigantea, and 
Puccinia Fuchsiae, shows resemblance to Puccinia Jmsiaeae. The 
second group, represented by Puccinia Epilobii and Puccinia scandica, 
shows a relationship to Puccinia Veratri and to Puccinia Epilohii- 
tetragoni. The third, represented by Puccinia sphaeroidea, is dis- 
tinctive, but evidently does not in reality belong among the rusts of the 
Onagraceae. 
II. Puccinia Circaeae Pers. Tent. Disp. Fung. 39. 1797. 
Literature: Saccardo, Syll. Fung. 7: 686. 1888. Schroeter, Pilz. Schl. i: 
348. 1889. Sydow, Monogr. Ured. i: 422. 1903. Fischer, Beitr. Krypt. Schweiz 
2^: 319. 1904. Holway, N. Amer. Ured. i: 79. 1907. Klebahn, Krypt. Mark 
Brand. 5^: 552. 1914. 
This cosmopolitan species occurs upon all the species of Circaea 
present in North America, the rust probably being coextensive with 
the host. It frequently has been noted that the tehospores germinate 
both as a micro- and a lepto-Puccinia, depending upon the season. 
The teliospores in this species are shorter and narrower than those 
of Puccinia Jussiaeae, yet it would seem that these are correlated 
species. The micro-form bears a greater resemblance to Puccinia 
Jussiaeae than does the lepto-form. 
Schweinitz, Schr. Nat. Ges. Leipzig i: 70. 1822, Wsted U redo 
Circaeae as occurring in Carolina, then in Trans. Amer. Phil. Soc. 
n. ser. 4: 291. 1832, gave the name as Caeoma Uredo Onagrarum 
Link, and Pennsylvania also as a locality. The names he gives are 
now considered to refer to Pucciniastrum Circaeae, a species which 
does not occur, so far as is known, in North America. In both the 
above mentioned publications Schweinitz also lists Puccinia Circaeae. 
Further information is being sought from the Schweinitz herbarium 
in Philadelphia; until an examination of the original material is made, 
if such a thing be possible, it may not be unreasonable to assume that 
