248 071 Ttiheritamce of Coat-Colour in the Greijhound 
According to some authorities black and red are the original greyhound 
colours ; white is said not to be found in any " natural breed of dogs." Black, red 
and white are often looked upon, however, as the three primitive " colours " by 
breeders. A mixture of these colours may then lead to a blend as in " fawn " and 
" blue," to particolour in large patches, to ticking in small spots or to stripes. 
According to Stonehenge we have the following pedigrees : 
(a) Bk. X W. 
^ 1 ^ 
Bk. W. W. Bk. Be. 
(b) Bk. X R. 
^ ^ ^ L-_ ^ 
Bk. Bd. R. R. Bk. Bk. Bd. Bk.' F. 
(c) R. X W. 
• ■ W. R. R. W. F. 
We are not convinced that such results pay sufficient attention to ancestry 
beyond the parents. Bd. x Bd. breeds as large a proportion true to parents as 
R. X R. or Bk. x Bk. Bk. x Bk. can produce R. and also F. The accompanying 
tables give the distribution of parents in the case of 2384 dogs and 2200 bitches. 
Here, to keep the tables within manageable limits, white has not been treated 
as a separate group except in the case of pure whites. Thus, of the red sires 
between \ and \ are red and white ; of the brindle sires between \ and i are 
brindle and white ; of the fawn sires more than \ are fawn and white ; of the 
mixed black sires ^ are black and white and about \ blue ; the remaining \ being 
chiefly blue and white, with a sprinkling of ticked and other rather nondescript 
dogs. Red shades off into fawn so that quite a large class of sires between ^ to 
\ of the reds are described as red or fawn. Besides these we find a few isolated 
units classed as red and fawn, or red, fawn and white. As the tables stand, pure 
black contains dogs without mixture of red, fawn, brindle or white ; fawn contains 
fawn, white and fawn ; red contains red, red-fawn, red or fawn, and these with 
white. Brindle contains brindle, brindle-white with a few isolated units of 
brindle-reds and brindle-fawns. The blue-brindle and brindle-blacks are included 
in the mixed blacks. For more detailed classifications the reader must refer to the 
complete tables in the Appendix of this memoir. Now a reference to the above 
tables will show that melanism may appear in R. x R. crosses as well as in F. x F. 
crosses ; that Bd. x Bd. may produce pure white or pure black dogs ; that Bk. x R. 
may give white dogs ; and that pure black dogs may produce pure white, red or 
fawn dogs. To use Mendelian language, the whole race of greyhounds appears 
to be in a heterozygous condition, and such pedigrees as those we have quoted 
from Stonehenge appear to have little valency when laige numbers are studied. 
(3) White Dogs. One great defect in our data is made manifest by an 
examination of Tables I and II ; although we have there the parents of 4584 dogs 
