E. Y. Thomson, J. Bell and K. Pearson 53 
(4) On the relative Variability of the General Population and of the Nest 
Queens. Absolute Values. 
Table II gives the absolute variabilities of the 26 characters determined, and 
we at once observe most marked differences between the two series. 
TABLE 11. 
Standard Deviations of Characters for Nest and Popidation Queens. 
Character 
Population Qdken 
Nest Queen 
Ratio P. Q.jN. Q. 
Eight Wing 
Left Wing 
Ri^'ht Wing 
Left Wing 
Eight 
Wing 
Left 
Wing 
A 
B 
C 
D 
E 
F 
G 
-.3124 + -0109 
•2099 ±-0073 
■1644+ -0057 
•1,343+ -0047 
•0527 + ^001 8 
•0991 + ^0034 
•0495f0017 
-3165 + -01 10 
-2026 + -0070 
-1678 ±-0058 
-1438 + -0050 
■05.52 fOO 19 
•1044 + -0036 
•0524+ •OOlS 
•1428 + ^0060 
•1089 + ^0046 
•0666 + ^0028 
•0525 + ^0022 
•0329 + ^0014 
•0462 + ^0019 
•0.376 ±^001 6 
-1464+ -0061 
•11 74 + -0049 
-0704 + -0030 
•0528 + ^0022 
-0246 + -0010 
-0468 + -0020 
-0385 + -0016 
2^1877 
r9275 
2^4685 
2-5.581 
1 -6018 
2^1450 
1-3165 
2-1619 
1- 7257 
2- 38.35 
2-72.35 
2-2439 
2 2308 
r.3610 
Mean 
2-029 
2-119 
H 
I 
J 
K 
L 
M 
•00914 + •000.32 
•01022+ •000.36 
•01022 + ^00036 
•00614 + -00021 
■00531 + -00018 
-00487 + -00017 
-00925 +-000.32 
-01058+ -00027 
-00881 ±-00031 
•00616 + -00021 
-00540+ -00019 
•00483 + ^00017 
•00329 + -00014 
-00628 + -00026 
-00717 + -00030 
-00422 + -00018 
-00281 + -00012 
-00.360+ -00015 
-00314 + -0001 3 
-00526 + -00022 
-00733+ -00031 
-00.504+ -00021 
-00285 + -0001 2 
•00345 ±-00014 
2-7781 
1-6274 
1-4254 
1 -4550 
1 -8897 
1 -3528 
2-9459 
2-0114 
1-2019 
1 -2222 
1 -8947 
1 -4000 
Mean 
1-755 
1-779 
Confining our attention to the absolute variabilities, we see that the variability 
of the queens in a single nest is not quite half the variability of the queens of the 
general population. Dr Warren has recently compared the absolute variability for 
the head index of Natal Termites for the caste of small soldiers* in the two cases 
of the general population and the population of a single nest. He found for the 
general population small soldiers a standard deviation of 17'08, and for the average 
nest one of 7-37, giving a ratio of 2-32. This is of the same order as our results, 
2-02 to 2-12. But Dr Warren noticed that between November and March the 
nest variability could alter considerably (9*00 to 6-56). Further, he was dealing 
with a head index, and we have compared it with absolute measurements on the 
wing. If we consider our index measurements on the wing they give (lower part 
of Table II) 1-76 to 1'78, as against his 2 32. When we consider, however, that 
he has only taken a single character against our averages of 14 absolute and 
12 index characters, and that we get ratios above and below his value, it is difficult 
* Biometrika, Vol. vi. pp. 336 and 339. 
