COOPERATIVE INVESTIGATIONS ON PLANTS. 
III. ON INHERITANCE IN THE SHIRLEY POPPY*. 
SECOND MEMOIR. 
CONTENTS. 
Page 
(1) Material and Manner of Fertilisation 394 
(2) Nature of the Observations 398 
(3) Influence of Environment on tlie Shirley Characters 401 
(4) Influence of Selection of Maternal Plants ....... 403 
(5) Influence of Variability of like Organs — Homotyposis — On Coefficients of 
Heredity 409 
(6) Parental Correlation ............ 410 
(7) Fraternal Correlation . ' 413 
(8) General Conclusions ............ 415 
Appendix: Correlation Tables .......... 41(J 
(1) Material and Manner of Fertilisation. 
A first memoir on the siibject of inheritance in the Shirley Poppy was publLshed 
in November, 1902-f-. In summing up our results in that paper we stated that: 
" Most plant organs being multiple in appearance, we have to apply special 
methods to deduce the intensity of heredity from multiple observations on the 
individual. It will probably be better in future experiments to confine attention 
to the first or principal flower, instead of using the indirect method of homotyposis ; 
but this will involve the observation (previous to harvesting) of individual plants 
in large series — 500 to 1000 — and much increase the labour of superintendence 
and observation J." 
We continued : 
" Notwithstanding the difficulties referred to above we find that for a variety 
of plant characters in the Shirley Poppy the values of hereditary influence found 
are on the whole in fair agreement with the like values in man." 
* These experiments have been materially assisted by the Grant made to the Department of Applied 
Mathematics at University College, London, by the Worshipful Company of Drapers. The establish- 
ment of a paid calculating staff has in addition largely accelerated the labour of reduction. 
t Biometrika, Vol. ii. pp. 56-100. + Ibid., p. 83. 
