III. Inheritanm in Shiiiey Poppy 
407 
and thus mucli alike, and very close to the Parkstone mothers of the Crewe crop. 
It seems therefore impossible to assert that the maternal plants at Crewe, although 
certainly an " inverse " selection from the Parkstone crop, are really a selection 
from the general population of Shirleys. We can only consider them as an average 
sample. 
(e) The Basal Patch. With the sole exception of wrinkling the basal patch 
is the most unsatisfactory of the characters observed. Not only do the flowers 
showing no basal patch vary very much in percentage with the environment, but 
there is undoubtedly a considerable amount of divergence in the estimations made 
by different observers of the categories. We have the following results : 
Parkstone General Parkstone Mothers Crewe Crop 
Population of Crewe Crop 
n 
MS y= Slight or none 295 134 53 
sd 
d 
dl 
I 
= Definite ... 342 128 214 
= Large ... 332 137 132 
969 399 399 
Or, in percentages 
Parkstone General Parkstone Mothers Crewe Crop 
Population of Crewe Crop 
Slight or none 30-4 33-6 13-3 
Definite ... 35-3 32 1 53-6 
Large ... 34-3 34-3 33-1 
Thus we see that while the maternal plants of the Crewe crop were a quite 
reasonably random sample of the Parkstone population, the distribution of Crewe 
offspring is totally different. How far is this due to environment, and how far to 
personal equation ? The corresponding percentages at Kidderminster, Arncliffe, 
and Streatham are : 
Kidderminster General Arncliffe General Streatham General 
Population Population Population 
Slight or none 36-5 18-4 52-4 
Definite ... 50-4 47-7 30 '9 
Large ... 13-1 33-9 16-7 
If these greater differences are not due to personal equation, which series can 
be taken as a typical crop of Shirleys ? We confess that there does not seem 
to us any means of separating out the environmental influence from the personal 
equation in this case. All we can say is that there is no evidence that the 
maternal plants at Crewe were a selection from the general population of Shirleys, 
and if they were, we have no means of measuring the stringency of that selection. 
if) Wrinkling of Petals. This, which is a marked feature of the beauty of 
the Shirley, is the most unsuccessful of all the characters recorded. The instruction 
to observe it between 12 and 24 hours after opening of the bud could not always 
