ON THE INHERITANCE OF COAT-COLOUR 
IN CATTLE. 
PART I. SHORTHORN CROSSES AND PURE SHORTHORNS. 
By a. BAERINGTON and K. PEARSON. 
CONTENTS. 
Page 
(1) Introductory 427 
(2) Material 430 
(3) Colour Inheritance in Blue-Grey Cattle 431 
(4) Horn Inheritance in Blue-Grey Cattle ....... 435 
(5) Colour History of Shorthorns 436 
(6) Possible Results of Various Colour Matings and Mendelian Interpretations . 440 
(7) Assortative Mating in the Shorthorn Population 444 
(8) On Direct Inheritance. Fir.st Degree 445 
(9) On Direct Inheritance. Second and Higher Degrees 448 
(10) On Collateral Inheritance. Whole and Half Siblings .... 453 
(11) General Conclusions 454 
(1) Introductory. 
As far as we are aware no statistical investigation into the inheritance of coat- 
colour in cattle has hitherto been published. An attempt will be made in the 
present memoir to cover some of this ground in a manner similar to that in which 
investigations have already been made in the case of the coat-colours of the 
thoroughbred horse* and of the greyhoundf. It is, perhaps, idle to assert that we 
have throughout desired to give as complete and accurate an account as lay within 
our power of the mass of material available, and this without prejudice in favour 
of any theory. . Those acquainted with the spirit which animates the work of 
biometricians will scarcely doubt this ; those who have throughout misrepresented 
the purport of mathematical investigations into heredity will doubtless continue 
to do so. At the same time it may not be without service to reiterate what has 
* Phil. Trans. Vol. 195 A, pp. 92-102 ; Biometrika, Vol. ii. pp. 229-210. 
t Biometrika, Vol. ni. pp. 245-298. 
