Contrihutions to a Knoiuiedge of South African OligocJiacta — Part II. 245 
tozoa in various stages of development, enclosed in definite " sperm " sacs. 
Matm-e spermatozoa were found in the spermathecae. 
■ The funnel of the sperm duct lies in segment xi, and, as is so frequently 
the case in Phreodrilidae, there is attached to its anterior face a mass of 
material taking in part a deep stain . By some they have been considered 
to be spermatozoa, but in the specimens of this new form examined by us 
they are so clearly continuous with the cells of the funnel that we regard 
them as consisting of long cilia. The central portion of the mass, unlike 
the anterior and posterior portions, has taken an intense blue stain with 
haematoxylin, and this is probably due to some secretion. Further, the 
whole mass is so compact and the area of the deeply stained portion is so 
regular. The funnel itself is composed of large cells appearing in sections 
as oblong shaped, with a slightly elongate nucleus centrally situated. 
The spermiducal gland occupies the greater portion of segment xii. It 
is a large U-shaped structure, with its convexity directed ventrally. The 
anterior limb lies close behind the anterior septum of segment xii, its pos- 
terior limb immediately in front of the penial sac. Its wall is composed 
of large vacuolated cells with basally situated spherical nucleus. 
The posterior limb rises to about the height of the centre of the segment 
and then becomes strongly attenuated, but the cells of this attenuated 
portion do not differ from those of the wide portion of the duct except 
in size. 
The thin tubular end of the spermiducal gland is now joined by the 
sperm duct. The atrium thus formed is composed of flattened epithelial 
cells and almost immediately enters the penis. 
The penial sac communicates with the exterior towards the posterior 
region of xii. It is a very long structure reaching to the middle of the 
height of the segment. In connection with its upper extremity is a band 
of retractor muscle ; and a large mass of muscular material is attached to 
the middle of its anterior surface. 
Its wall is composed of the three sets of elements usually present. 
Its inner epithelium is composed of loosely arranged cells with slightly 
elongate nuclei. The outer layer is constituted by an indistinct peritoneal 
epithelium with flattened cells. It bears a very close resemblance to the 
corresponding structure in Gondwanaedrilus. 
The penis is peculiar in that it appears as a double tube. It is an 
elongate slender structure extending practically the whole length of the 
penial chamber. 
Its outer surface is constituted by a folded epithelium of squarish cells 
with spherical nuclei, and bearing a strong resemblance to the inner layer 
of the penial chamber. Beneath this epithelium is a basement membrane, 
and within this there is a clear and indistinct layer in which in places 
spherical nuclei can be detected. The continuation of the atrium passes 
