88 
Transactions of the Royal Society of South Africa. 
plantar side) is the intermedium. Lateral to this is the large fibiale with a 
pronounced heel. This structure differs from that of Howesia in the fusion 
of the tibiale and centrale. Such a fusion is paralleled in stage Q of 
Sphenoclon ; and a further fusion with the intermedium is seen in Procolo- 
phon. In the rest of the foot Hoivesia and Mesosuchus agree. A specimen 
from Burghersdorp (S.A. Mus. Cat. No. 5861) collected by Dr. du Toit 
shows the fingers and toes of a species of either Hoivesia or Mesosuchus. The 
fifth metatarsal is of exactly the same type as in those genera, and the 
digital formula of the pes is 2, 3, 4, 5, 3. The third metatarsal is the longest. 
The femur in the Howesiidae differs from that of the Pseudosuchia such 
as Euparlceria and approximates more closely to that of Erythrosuchus, but 
in the latter the large medial trochanter is not terminal. The forms agree 
with Erythrosuchus, too, in the sudden bending down of the anterior portion 
of the pubis ; but the ilium lacks the posterior prolongation seen in both 
Eryth rosu ch u s and Euparlceria . 
EXPLANATION OF PLATES II and III. 
1. Euparlceria capensis. 
2. Exiparlceria capensis. 
3. Euparlceria capensis. 
4. Euparlceria capensis. 
5. Euparlceria capensis. 
6. Broivniella africana. 
7. Euparheria capensis. 
8. Euparlceria capensis. 
PLATE II. 
Outer view of left dentary of Specimen No. 2. 
Lower view of same. 
Lateral view of humerus of Specimen No. 2. 
Medio-anterior view of same. 
Femur. 
Femur. 
Pelvis of Specimen No. 2. 
Right ilium of Specimen No. 3. 
PLATE III. 
Mesosuchus broxvni, Watson. 
1. Left ilium. 
2. Pelvis from below. 
3. Proximal end of femur. 
4. Femur. 
5. Fore limb, showing carpus. 
6. Hind limb, showing tarsus. 
7. True view of metatarsal V. 
