HOWARD HUGHES MEDICAL INSTITUTE 
SENIOR INVESTIGATORS, INVESTIGATORS, ASSOCIATE INVESTIGATORS, 
AND ASSISTANT INVESTIGATORS 
SEPTEMBER 1, 1991 -AUGUST 31, 1992 
SENIOR INVESTIGATORS 
Kandel, Eric R., M.D. 
Leder, Philip, M.D. 
Nathans, Daniel, M.D. 
Steiner, Donald F., M.D. 
INVESTIGATORS 
Adams, Paul R., Ph.D. 
Alt, Frederick W., Ph.D. 
Artavanis-Tsakonas, Spyridon, Ph.D. 
Atkinson, John P., M.D. 
Axel, Richard, M.D. 
Beach, David H., Ph.D. 
Beaudet, Arthur L., M.D. 
Bell, Graeme I., Ph.D. 
Bennett, G. Vann, M.D., Ph.D. 
Bevan, Michael J., Ph.D. 
Blackshear, Perry J., M.D., D.Phil. 
Blobel, Gunter, M.D., Ph.D. 
Bloom, Barry R., Ph.D. 
Brugge, Joan S., Ph.D. 
Campbell, Kevin P., Ph.D. 
Capecchi, Mario R., Ph.D. 
Caron, Marc G., Ph.D. 
Caskey, C. Thomas, M.D. 
Cech, Thomas R , Ph.D. 
Chin, William W., M.D. 
Collins, Francis S., M.D., Ph.D. 
Cooper, Max D., M.D. 
Cresswell, Peter, Ph.D. 
Davis, Mark M., Ph.D. 
De Camilli, Pietro, M.D. 
Deisenhofer, Johann, Ph.D. 
Donelson, John E., Ph.D. 
Dreyfuss, Gideon, Ph.D. 
Esmon, Charles T., Ph.D. 
Evans, Ronald M., Ph.D. 
Exton, John H., M.D., Ph.D. 
Fischer Lindahl, Kirsten, Ph.D. 
Flavell, Richard A., Ph.D. 
Francke, Uta, M.D. 
Fuchs, Elaine, Ph.D. 
Garbers, David L., Ph.D. 
Gesteland, Raymond F., Ph.D. 
Gething, Mary-Jane H., Ph.D. 
Glomset, John A., M.D. 
Goodman, Corey S., Ph.D. 
Greene, Warner C, M.D., Ph.D. 
Habener, Joel F., M.D. 
Harrison, Stephen C, Ph.D. 
Heintz, Nathaniel, Ph.D. 
Hendrickson, Wayne A., Ph.D. 
Henikoff, Steven, Ph.D. 
Horvitz, H. Robert, Ph.D. 
Hyiies, Richard C, Ph.D. 
Jan, Lily Y., Ph.D. 
Jan, Yuh Nung, Ph.D. 
Janeway, Charles A., Jr., M.D. 
Jessell, Thomas M., Ph.D. 
Kaback, H. Ronald, M.D. 
Kan, Yuet Wai, M.D. 
Kappler, John W., Ph.D. 
Kaufman, Thomas C, Ph.D. 
Kunkel, Louis M., Ph.D. 
Kurnit, David M., M.D., Ph.D. 
Lai, Michael M.-C, M.D., Ph.D. 
Lalouel, Jean-Marc, M.D., D.Sc. 
Lamb, Robert A., Ph.D., Sc.D. 
Larsen, P. Reed, M.D. 
Lefkowitz, Robert J., M.D. 
Liebhaber, Stephen A., M.D. 
Lindquist, Susan L., Ph.D. 
Mailer, James L., Ph.D. 
Marrack, Philippa, Ph.D. 
Massague, Joan, Ph.D. 
Matthews, Brian W., Ph.D., D.Sc. 
McKnight, Steven L., Ph.D. 
Miller, Christopher, Ph.D. 
Movshon, J. Anthony, Ph.D. 
Nadal-Ginard, Bernardo, M.D., Ph.D. 
Nevins, Joseph R., Ph.D. 
Olson, Maynard V., Ph.D. 
Orkin, Stuart H , M.D. 
Pabo, Carl O , Ph.D. 
Palmiter, Richard D., Ph.D. 
Perlmutter, Roger M., M.D., Ph.D. 
Quiocho, Florante A., Ph.D. 
Reichardt, Louis F., Ph.D. 
Rosbash, Michael, Ph.D. 
Rosenfeld, Michael G., M.D. 
Rubin, Gerald M., Ph.D. 
Schekman, Randy W., Ph.D. 
Schw^artz, James H., M.D., Ph.D. 
Sedat,John W., Ph.D. 
Seidman, Jonathan G., Ph.D. 
Sejnowski, TerrenceJ., Ph.D. 
Shenk, Thomas E., Ph.D. 
Sherr, Charles J., M.D., Ph.D. 
