370 
FERTILISATION OF THE GOODEN1ACE.V, 
however, deeply coloured. The colour is very deep purple. The 
indusium is two -lipped, and in mature flowers closes, except for a 
small central opening. In all the older flowers I examined the 
indusium was full of pollen, and the auricles lined with a sheet of 
adhering grains. The indusium and stigma are purple, the style 
below green. In no instance did I see the stigma outgrown, 
or even so near the opening as to be capable of receiving pollen 
from a visiting insect, so that this species also falls within the 
group not fully adapted for insect-fertilisation; this is therefore 
another species with arrangements for fertilisation complete 
except at one point. 
Summing up, it appears that in Dampiera there is a complex 
mechanism directed towards the accomplishment of cross-fertilisa- 
tion by insects, and yet most species examined stop short of 
completeness, from the stigma's not growing out so as to be exposed 
to the touch of pollen-laden visitors. Those examined may be 
divided into two groups; those having either a shallow indusium 
so that the stigma may be reached by insects; and those in which 
the indusium is deep, and from the stigma's not growing out, 
incapable of being insect-fertilised. In the first of these groups 
are D. linearis, sp. (?), eriocephala and juncea; in the latter D. 
Brownii, stricta, luteiflora, Linschotenii, loranthifolia, laneeolata, 
and adpressa. 
The various species I have examined show a gradation in the 
completeness of adaptation of the various parts. Thus in the 
auricles there is a progression from the simple fold with a slight 
hollow in the centre (D. linearis) to the most complex arrange- 
ment of folds, hairs and trichomes as in I), stricta and D. Brownii. 
And in those species which show this gradual increase of adapta- 
tion, there is also a regular augmentation of colour in the auricles, 
from the simplest with a patch of faint colour in the centre, to 
the deep purple spread all over the auricle in the most complex 
forms. This is also the case in the style, which varies from green 
to purple, and the indusium, from pale red to purple. The stigma 
is always coloured, in which the genus differs from almost every 
other member of the order. The indusium also varies from the 
