I 3 
MORPHOLOGY 
is observed throughout the groups having archegonia. The primary 
ventral cell, just before the maturity of the archegonium, divides into 
the ventral canal cell and the egg, and finally all the canal cells dis- 
organize (fig. 287). 
287 
FIGS. 283-287. Archegonium of Lycopodium clavatum: 283, young archegonium 
after the first division (periclinal) of the superficial initial, resulting in the primary 
neck cell (outer) and inner cell; 284, division of neck cell and also of inner cell, the 
latter division resulting in basal cell and central cell (both shaded); 285, division of 
central cell, giving rise to primary neck canal cell (outermost shaded one) and pri- 
mary ventral cell (middle shaded one); 286, the completed axial row, consisting of six 
neck canal cells, the ventral canal cell, and the egg; 287, breaking down of canal cells, 
leaving a passageway to the egg. After BRUCHMANN. 
Embryo. The embryo of the Lycopodium sporophyte develops in 
a very characteristic way (figs. 288-293). The fertilized egg divides 
by a transverse wall, the resulting outer cell being the suspensor cell, 
and the inner one the embryonal cell (fig. 288). The suspensor cell 
may or may not divide, but in any event it usually becomes elongated. 
The suspensor is an organ of the embryo, but does not enter into its 
permanent structure, which is developed by the embryonal cell. By 
successive divisions this cell becomes four cells (figs. 289, 290), which 
are related to each other as unequal quadrants of a sphere. Two of 
these quadrant cells develop the foot, which is an absorbing organ of 
the embryo while it is feeding upon the gametophyte. One of the 
remaining quadrant cells develops the stem, and the other the first leaf. 
It is to be observed that in this first differentiation of body regions the 
root is omitted, but it develops later from the tissue produced by the 
leaf quadrant (fig. 294). The foot is a structure of the sporophyte 
found among the bryophytes, among which the sporophyte is dependent 
on the gametophyte through life. The suspensor is a new organ of the 
