222 
MORPHOLOGY 
the stamen is peltate (as in taxads), and one-sided peltate (as in Tor- 
reya). The sporangia are developed as usual, and in the Abietineae the 
pollen grains (microspores) are winged (as in podocarps, fig. 489). 
Ovulate strobilus. The ovulate strobilus of Pinaceae has been the 
subject of much discussion. In the Abietineae the strobilus is made 
up of a series of bracts, in the axil of each of which a so-called ovulif- 
erous scale appears, which usually bears two ovules whose micropyles 
are directed towards the base of the scale (fig. 490). In the other tribes 
the bract and ovuliferous scale are more or less united. The discussion 
492 
FlGS. 491, 492. Archegonium complex of Thuja: 491, group of archegonium 
initials; 492, two mature archegonia (reached by a pollen tube, in which the body cell 
has not yet divided) with a common archegonial chamber and a common archegonial 
(nutritive) jacket. After LAND. 
referred to has to do with the nature of the ovuliferous scale, and many 
facts indicate that it represents a fused pair of leaves of a dwarf axillary 
branch. This means that the ovules are borne in the strobilus on axes 
of the second order, as in Cordaitales, and that the ovulate strobilus of 
Pinaceae is a compound strobilus. 
Ovule. The structure of the ovule is as described for Taxaceae, 
except that the outer fleshy layer does not develop, and the seed is dry ; 
that both sets of vascular strands have been eliminated ; and also that 
there is the same elaborate nutritive mechanism that was described 
for Ginkgo. The development of the ovule and the seed is usually 
