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Research Ballet hi No. 4 
the development of the pigment in order to produce a pat- 
tern at all, they are now known to be distinct in inherit- 
ance from the factors for pigment — a fact that I have 
been able to show by use of a race of maize with a peculiar 
brown pericarp in addition to races with red pericarp. 
Summaky 
A somatic variation in maize is shown to be inherited in 
simple Mendelian fashion. The variation has to do with 
the development of a dark red pigment (or in one stock 
a brown pigment) in the pericarp of the grains, often 
associated with the development of an apparently similar 
pigment in the cob and husks. 
Plants in which this pigment has a variegated pattern 
may show any amount of red pericarp, including wholly 
self-red ears, large or small patches of self-red grains, 
scattered self-red grains, grains with a single stripe of 
red covering from perhaps nine tenths to one tenth of the 
surface, grains with several prominent stripes and those 
with a single minute streak, ears with most of the grains 
prominently striped and ears that are non-colored except 
for a single partly colored grain, and probably also plants 
with wholly self-red and others with wholly colorless ears. 
It is shown that the amount of pigment developed in the 
pericarp of variegated seeds bears a definite relation to 
the development of color in the progeny of such seeds. 
This relation is not such that seeds showing say nine 
tenths, one half, or one tenth red will produce or even tend 
to produce plants whose ears as a whole or whose indi- 
vidual grains are, respectively, nine tenths, one half, or 
one tenth red. Experimental results indicate rather that 
the more color in the pericarp of the seeds planted the 
more likely are they to produce plants with wholly self- 
red ears, and, correspondingly, the less likely to yield 
plants with variegated ears. 
Self -red ears thus produced are shown to behave in in- 
