284 
HICK : CALAMOSTACHYS BINNEYANA. 
In fig. 1, PL L, these canals are well and distinctly shown at «, 
but they appear to have been enlarged by the tearing of the peri- 
pheral tissue, as they are smaller in all the other sections. At the 
margin we observe small elements, v, somewhat different from the 
rest, projecting into the open space, which forcibly remind us of what 
is found in the carinal canals of Equisetum. This and other similar 
sections are of themselves almost conclusive of the nature of these 
open spaces, but the longitudinal one shown in figs. 2, PI. I., and 
3, PL II., is absolutely demonstrative. Here we have one of the 
canals cut longitudinally, c, and at various points along its course we 
find, adhering to its sides, fragments of annular, and spiral vessels 
almost exactly as they are seen in longitudinal sections of the steins 
of living Equiseta, fig, 3, v. PL II. There are remnants too of the 
tissue which originally interrupted the continuity of the canals at 
the " nodal disks," one of which is shown at d. 
I am convinced then that we have here structures which are 
strictly homologous with the carinal canals of Equisetum and the 
extinct Arthropitys. Taking the view of Count Sohns, that in these 
two genera the canals represent the initial tracheal strands of vas- 
cular bundles, we reach the conclusion already stated, that in the.^e 
specimens of Codamostachys Binneyana the axis not only had a 
parenchymatous pith, but that the primary vascular bundles were of 
the Calamitean type. 
An examination of numerous other sections, while confirming 
these conclusions, reveals one or two points of difference which it 
may be well to mention. In some the pith has a triangular rather 
than a circular section, and in these cases the carinal canals are 
situated at the angles. The angles however are usually truncated, 
and are not always of the same breadth tangentially. Sometimes 
indeed they give the impression that they are about to bifurcate, the 
carinal canals dividing, so to speak, at the same time. Obviously 
we have here an increase in the number of the primary bundles, and 
in this way the number of canals may be increased to six, a number 
actually found in two sections belonging to Mr. Lomax, of Radcliffe, 
which he permitted me to examine. In another case, five transverse 
sections have been cut from the same strobilus, four of which have 
