288 
hick: calamostachys BINNEYANA. 
tology is relatively simple. Cellular in the main, the peripheral 
tissue of the pedicel, is apparently sclerenchymatous, while a vascular 
strand runs up the center. The sclerenchyma is continued on the 
underside of the scutellum, while the vascular .strand divides into 
four .smaller strands, which diverge to the four sporangia. The 
conical mass of tissue which lies between these diverging strands and 
the external surface of the scutellum appears to have been of a deli- 
cate nature, as it is frequently wanting when the other tissues are 
well preserved. The superficial cells of the external surface consti- 
tute a well-developed epidermis whose elements are large and 
elongated vertically to the surface, but their walls are not thickened 
as in Equisetum. This last character is no doubt to be correlated 
with the fact that they are protected by the bracts from the nodal 
disk below. 
It does not appear to have been previously made known that 
the annular or spiral trachea? which form the vascular strands of the 
sporangiophores are accompanied by elongated, thin- walled, narrow 
cells, which have some of the appearances of a phloem tissue. This 
is especially evident where the vascular tissue approaches the point 
of attachment of the sporangia. So distinct and perfect are these 
cells at this point in one of the sections, that in some of them struc- 
tures are seen that almost compel the belief that they are nuclei ! 
Professor Williamson has shown* that the sporangia are not 
connected with the scutellum by the entire base as in Equisetum, but 
by a narrow neck of cellular tissue attached a little within the over- 
hanging margin. One of these peduncles is seen in fig. o, PI. II. 
This mode of attachment of the sporangia seems to be an important 
feature of this species of Galamostachys, as it appear.- to indicate 
that the archesporia were developed in special lobes of the scutellum. 
The Sporangia. 
The form and structure of the sporangia has been 60 well des- 
cribed already that they need not be dealt with here. One feature 
however may be mentioned as it does not seem to have been 
previously brought forward, or at all events sufficiently emphasised. 
Op. cit, Ft. xi. 5 1881, p. 298. 
