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MR.  R.  OWEN  ON  THE  ANATOMY  OF  THE  CHEETAH. 
In  the  description  of  the  outward  configuration  of  the  cerebral  hemispheres  in  the 
Cheetah  and  other  feUne  species,  I  have  hmited  myself  to  noting  those  convolutions  only 
which,  after  a  careful  comparison  of  the  materials  at  my  disposal,  appeared  to  be  sub- 
ject to  least  variety.  But  even  with  this  limitation  a  very  small  portion  of  the  cerebral 
surface  remains  undescribed ;  and  the  constancy  manifested  in  the  disposition  of  the 
remainder,  as  to  the  form,  extent,  and  symmetrical  arrangement  of  the  convolutions, 
argues  strongly  in  favour  of  the  conclusion  that  the  folding  of  the  hemispheric  sub- 
stance in  the  progress  of  its  development,  follows  a  determinate  law ;  and  that  the 
tracing  of  the  additional  convolutions,  as  they  successively  present  themselves  in  suc- 
ceeding complexities  of  the  cerebrum,  may  not  only  tend  to  advance  zoology  by  bringing 
to  light  additional  instances  of  affinities  between  the  different  groups  of  Mammalia,  but 
ultimately  lead  to  the  determination  both  of  the  amount  and  locality  of  the  convo- 
lutions in  the  human  brain  which  are  analogous  to  those  of  the  inferior  animals. 
PLATE  XX. 
Fig.  1 .  Superior  view  of  the  brain  of  the  Cheetah. 
2.  Side  view  of  the  same. 
3.  Mesial  surface  of  the  right  hemisphere  of  the  same. 
4.  Superior  view  of  the  brain  of  the  domestic  Cat. 
5.  Side  view  of  the  same. 
6.  Mesial  surface  of  the  right  hemisphere  of  the  same. 
A.  The  cerebrum ;  B.  the  cerebellum ;  C.  the  medulla  oblongata ;  D.  the  spinal  cord. 
The  smaller  letters  and  figures  are  explained  in  the  text. 
anteriorly,  and  forms  a  large  portion  of  the  anterior  lobe :  the  middle  one  turns  outwardly,  and  joins  particu- 
larly the  external  lateral  mass,  which  does  not  extend  farther  forwards  than  about  two  thirds  of  the  whole 
extent  of  the  cerebrum  :  the  external  or  lateral  mass  is  subdivided  by  two  transverse  perpendicular  fissures  into 
three  convolutions,  of  which  probably  the  posterior  may  be  Combativeness,  the  middle  Destructiveness,  the 
anterior  Secretiveness  and  Alimentiveness ;  these  three  all  unite  below. 
"  The  under  surface  of  the  anterior  lobe  is  divided  by  a  fissure  extending  nearly  in  the  direction  of  the  outer 
margin  of  the  olfactory  nerve,  as  in  Man,  in  whom  the  mesial  convolution  contains  Individuality :  this  in  the 
Cat  may  perhaps  include  other  organs. 
"  The  brains  of  the  whole  genus  Felis  are  similar  as  to  these  general  divisions,  though  the  convolutions  vary 
as  to  their  relative  proportions  in  each  species,  and  frequently  in  individuals  of  the  same  species. 
"  In  comparing  the  genus  Felis  with  the  Dog  tribes,  the  posterior  internal  longitudinal  mass  is  much  smaller 
than  the  middle ;  and  in  the  Jackal  the  middle  mass  is  half  as  much  more  voluminous  as  the  internal  mass, 
while  in  most  of  the  Cats  these  parts  are  nearly  equal,  and  in  some  the  internal  preponderates.  The  posterior 
division  of  the  external  lateral  mass,  Combativeness,  is  smaller  than  the  middle  one,  Destructiveness,  in  the 
Cats,  while  the  opposite  fact  appears  in  the  Dogs :  in  this  respect  the  Lion  approaches  more  to  the  Dog  tribe 
than  any  of  the  genus  Felis." — H.  H.  H. 
