YOSHIMARO TANAKA 
141 
The fact that deserves notice in the above results, is the total absence of 
distinct quails which were to be expected if the Q, factor is independently 
transmissible. On the other hand, the Spq form, if present, ought to be an 
apparent combination of S and pq patterns, namely a very light form of SpQ,; 
but no individuals which could be regarded as "striped-pale-quail" appeared 
at all. Hence it follows that Q, is completely coupled with S in both sexes. 
Moreover, the linkage is somewhat peculiar in that S can never exist without 
Q, while Q, can be present regardless of the S factor. A similar relation is 
advocated by Punnett (1915) for D and E factors in the coat colour of the 
rabbit. 
The following data are explicable on the same assumption. — 
Triple hybrids were produced by crossing the striped (SPQ,) with the pale- 
quail (spq); the hybrids were either inbred or back-crossed with the re- 
cessives. 
Triheterozygous striped (SsPpQq) inbred. 
Striped 
(SPQ) 
Striped-quail 
(SpQ) 
Plain 
(sPq) 
Pale-quail 
( s pq) 
Totals 
A 52i'i5 
119 
53 
42 
18 
232 
A 524' j 5 
122 
64 
46 
18 
250 
A 53i'i5 
76 
5' 
38 
2 1 
186 
A 532'i5 
' '3 
40 
41 * 
10* 
204 
A 539/15 
224 
85 
72* 
28 * 
409 
A 540' 1 5 
1 20 
46 
40 
24 
230 
Totals 
774 
339 
279 
119 
1511 
Expect. 
850 
283 
283 
95 
1511 
Ratio 
9 
: 3 = 
3 : 
1 
Triheterozy 
gous striped (SsPpQq) 2 2 
x pale-quail (ssppc[q) J J. 
Striped 
Striped-quail 
Plain 
Pale-quail 
Totals 
A 5 3 6'i 5 
58 
74 
65 
95* 
292 
A 537'<5 
80 
85 
78* 
84* 
327 
Totals 
138 
'59 
H3 
179 
619 
Expect. 
'54-75 
'54-75 
'54-75 
' 54-75 
619.00 
