YOSHIMARO TANAKA 145 
other marking but P (PqL), multilunar spots become so faintly yellowish 
that one might overlook them unless special attention is given, whereas 
in multilunar-normal (PQL), multilunar-quail (pQL), and even in multilnnar- 
pale-quail (pqL), the spots under consideration are much more distinct. Conse- 
quently there are cases apparently peculiar, in which PqL animals heterozygous 
for P give more deeply pigmented multilunar offspring (pqL), when inbred or 
cross-bred with recessive. On the other hand, Z develops the characteristic 
black bands which are nearly equally intense without regard to the presence 
or absence of P or of other marking factors. 
III. MULTIPLE ALLELOMORPHISM AND COMPLETE 
REPULSION. 
The inheritance of markings, which was looked upon by Sturtevant 
(19 1 5) as being due to multiple allelomorphism, has been proved in the majority 
of cases to be owing to complete coupling, as explained in the preceding 
chapter. There is, however, a single case which must be explained on the 
basis of complete repulsion. This is the subject of this section. 
It was mentioned in a paper of mine (1914a, p. 6) that a striped 0 (ex 
striped x moricaud) yielded, by crossing with the quail (Kasuri), no other 
recessives but striped and moricaud. This was the only case which suggest- 
ed a hereditary relation between S and M; but further data have since been 
obtained, so that we are justified in drawing the following conclusion. 
Fj individuals (SsMm) ex the cross striped (Sm) x moricaud (sM) appear- 
ed at first sight just the same as the striped parent. On careful examination 
it was found that the SM individuals distinctly differed from the ordinary 
striped ones (Sm) ; the former being provided, in addition to the S pattern, 
with the characteristic spots on the head-case, on the ventral side of the 
thorax, and on the inter-segmental bands, so that we are able to distinguish 
externally the Sm individuals from the SM, which will be designated as the 
"striped-moricaud." 
1) In reality striped-moricaud. 
