YüSHLMARO TAN AK A !8c 
while Emerson (191 1) made out a similar case in the colour correlation (com- 
plete coupling) of the maize. On the other hand, there are some authors 
who merely assumes the similarity of male and female gametic series without 
actually bringing them to the test mentioned above. 
A few previous investigators maintain, however, dissimilar gametic distri- 
bution, in the male and female, of the factors that are not sex-limited in 
several species of plants and one of animals. 
De Vries(io,ii, 191 3) discovered Oenothera hybrids which were pro- 
duced by reciprocal crossings and markedly different from each other, being 
apparently patroclinous or matroclinous ; he says : 
"Hier werden von den sichtbaren Eigenschaften einige nur mittels des 
Pollens und nicht durch die Eizellen auf die Kinder vererbt. Demgegen- 
über gibt es andere Merkmale welche nur in den Eizellen, und nicht im 
Pollen übertragen werden. Oder allgemein : Im Pollen und in den 
Eizellen werden verschiedene erbliche Eigenschaften auf die Nach- 
kommen übermittelt." 
"Heterogamy" as this phenomenon is called by de Vkies, or "allogamy" 
by Swingle (1911), was most striking, among various cases, in the crossing 
between 0. biennis and 0. muricata. 
Allogamy in Oenothera is regarded by Goedschmidt (1913a) from the 
cytological point of view as due to merogony, but Renner (191 3) rejects 
this view. 
Gregory (1911a) obtained the ratio 411 (magenta, green stigma) 198 
(magenta, red stigma) : 97 (red, green stigma) : 78 (red, red stigma) in Primula, 
and remarks as follows : 
"For the time being it may be pointed out that a very close approxi- 
mation to the observed numbers is given by the assumption that a 
coupling of the form 7 : 1 :i :j is present in gametes of one sex only, gametes 
of the opposite sex being produced in equal numbers of all four kinds."' 
Gregory's view is criticized by Trow (1912), who maintains that the 
ratio obtained may be expected on the basis of 2 : 1 : i : 2 coupling 
