112 
FUNGI. 
has noted the variability of the spores in Triphragmium,* and 
Patouillardf has some remarks bearing on the same subject in his 
description of T. setulosum. 
Three forms of spore, belonging to the teleutospore stage, are 
to be met with in every species, as follows : — (i.) The generally 
understood Triphragmium type, a radiately three-celled spore. 
The basal cell is seated on a more or less elongated pedicel ; each 
cell has a single germ-spore. This type is by far most abundant 
in every species (excepting T. acacice , Cke.), and in the accom- 
panying figures is indicated by a small x . (ii.) A two-celled 
spore, having the cells superposed and separated by a transverse 
septum, resembling the spore of a Puccinia ; each cell having one 
lateral germ-pore ; pedicel as in the first type, (iii.) A one-celled 
spore, with a single lateral germ-pore and basal pedicel. 
Variations of the types described above are common. In (1) 
the three cells are normally, that is most frequently, about equal 
in size, but, as the figures show, this state of things is by no means 
constant. Sometimes one or other of the cells is much reduced in 
size, and when very small is not furnished with a germ-pore. Not 
unfrequently there are two basal cells, divided by a vertical septum, 
the pedicel at first sight appearing to be fixed to the septum, and 
thus common to the two cells, but in every example of such that I 
have examined the pedicel is found to be organically united to one 
cell only, as shown in Fig. 4 x . 
Dietel states that in most species of Triphragmium the germ- 
pore perforates the inner layer of the endospore only.J 
The germination of the component cells of the teleutospore (in 
( T. ulmariae ) is basipetal, the pro-mycelium of the basal cell being 
formed last, and is as a rule not so robust as the tubes of the 
superior cells, and in not a few instances is not developed at all, 
or remains in a rudimentary condition. 
Magnus§ favours the view that the uredospores in the Uredineae 
have developed out of teleutospores, and the variability in the last- 
named type of spore in the genus Triphragmium appears to favour 
this view. As a rule the teleutospores are most constant in form 
in those species of the Uredineae where uredospores are also pre- 
sent. 
The accompanying figures belong to the following species, as 
understood in Saccardo’s “ Sylloge Fungorum,” Vol. vii., Pt. II. 
Fig. 1. — Triphragmium isopyri, Moug. (drawn from Mong. & Nest., Stirp. 
Cr., No. 892). 
„ 2. — T. ulmarice , Link, (drawn from Cooke’s Fung. Brit. Exs., No. 23). 
,, 3. — T. JilipendulcB , Pass, (drawn from Bab., Fung. Eur., No. 2082, 
Com. Passer.). 
* “ Ber. der Deutschen Bot. Gesellsch.,” 1891, p. 120 (1 pi.). 
f u Jonrn. Bot.” (Morot), 1890, p. 58, Fig. 3. 
% “ Flora,” lxxiv. (1891), p. 140 (1 pi.). 
§ “ Ber. Deutsch. Bot. Gesell.,’’ ix. (1891), Gen.-Versamml. Heft,, pp. 85 
(1 ph). 
