1891-92.] Prof. Sir Wm. Turner on the Lesser Rorqual. 55 
dilatation consequently as a pouch-like commencement of the 
duodenum. It is well known that in the common Porpoise and 
in Lagenorynclius alhirostris the duodenum begins in a suhglohular 
dilatation, into which the biliary and pancreatic ducts open. I have 
figured a corresponding dilatation in the stomachs of Delpliinus 
delpliis and Grampus griseus,^ which also receives the ducts of the 
liver and pancreas. If these ducts in B. rostrata opened into it, 
obviously it would be the duodenum; hut, unfortunately, in taking my 
specimen out of the abdomen the biliary and pancreatic ducts were 
not dissected out and preserved, so that I am unable to say from 
personal observation where they opened. On this point, however, 
Drs Carte and Macalister have made a definite statement, for not 
only do they call the dilatation a 5th compartment, but they say 
that it communicated with the duodenum by a small pylorus, and 
that the conjoined hepatico-pancreatic ducts, after running obliquely 
for 2 inches between the coats of the intestine, opened in a valve- 
like manner into the interior of the duodenum about 6f inches 
below the pyloric orifice. Eelying on the accuracy of this observa- 
tion, the dilatation should be regarded as a 5th compartment of the 
stomach. 
Something more is, however, recjuired in the study of the Cetacean 
stomach than the determination of the number and arrangement of 
the compartments : the structure of the mucous membrane has to 
be determined. In a memoir on Sowerby’s whale, published 
in 1885,1 I showed, from the examination of its stomach and that 
of the dried stomach of Hyperoodon, that in these animals, and 
presumably in the Ziphioid whales generally, the 1st compartment 
of the stomach is not a mere paunch-like receptacle lined by a pro- 
longation of the oesophageal epithelium, as in the Dolphins. It is, 
on the other hand, a true digestive chamber, the mucous membrane 
lining which is abundantly provided with gastric glands, like those 
of the 2nd compartment in the stomach of the Dolphins, with which 
it is therefore homologous. Shortly after the publication of that 
paper. Professor Max Weber came to the same conclusion from the 
study of the stomach of Hyperoodon. | More recently I have 
* Jour, of Anat and Phys., vols. xxiii. p. 479, and xxvi. p. 260. 
t Ibid., vol. XX. 
X Studien uber Sdugethiere, Jena, 1886. 
