1915-16.] Obituary Notices. 335 
fresh ground, and had provided a terminology which was in advance of 
the observed facts. What was then urgently required was cool and con- 
trolled observation : and this was exactly what G Wynne- Vaughan was 
so well fitted to supply. 
He first engaged in the examination of the stelar conditions seen in 
the Nymphaeaceae and the Primulaceae, and the results were published in 
1897 ; the former in a Memoir in the Transactions of the Linncean Society 
(“ On the Morphology and Anatomy of the Nymphaeaceae ”), the latter 
in the Annals of Botany (“ On Polystely in the genus Primula ”). At the 
British Association Meeting at Liverpool in 1896 he gave a preliminary 
account of his observations, which showed his hearers that not only a new 
investigator, but also a new teacher had appeared. It led to his appoint- 
ment as Assistant in the Botanical Department of Glasgow University. 
Lang was already a member of the staff there, and for ten years these two, 
with the Professor, worked together, each in his several way, but with 
full mutual knowledge, upon problems relating to the Pteridophyta. 
It is not always that Scottish students take to a teacher having 
marked racial characters not their own. But Gwynne- Vaughan, though 
a Welshman through and through, was a success in Glasgow from the 
first, with the large medical classes as well as with the smaller classes 
for women at Queen Margaret College. He must have handled some 
1500 students while in Glasgow, and it was clear that they appreciated 
the energy and single-mindedness of his work with them. On ex- 
cursions his activity, combined with his sporting instincts and wide 
natural knowledge, secured for him an enthusiastic personal following. 
He also took his share in the advanced teaching, laying in certain re- 
stricted branches the foundation of those valuable notes which were ex- 
panded over the whole area of the science during his later period of 
teaching in London. 
The anatomical experience which Gwynne- Vaughan had gained at Kew, 
and his grasp of stelar questions as illustrated in flowering plants, fitted him 
to enter with special insight on his arrival in Glasgow into the investi- 
gation of the anatomy of the Filicales. Mr Boodle was already engaged 
at Kew in similar work. But by mutual consent it was arranged that 
while he undertook more especially the Schizseacese, Gleicheniacese, and 
Hymenophyllacem, Gwynne- Vaughan should devote himself to types 
which showed greater advance in anatomical complexity. Accepting the 
protostelic state as probably the primitive condition for all, it became 
necessary by patient observation and comparison to relate to it the more 
complex states already recognised by Van Tieghem as “ polystelic.” 
