1915-16.] Obituary Notices. 337 
In relation to this work on the Fossil Osmundacese thFsre was also 
published a short but important joint note “ On the Origin of the Adaxi- 
ally-curved Leaf-trace in the Filicales” ( Proc . Roy. Soc. Edin., 1908, p. 433). 
There are also two later papers by G Wynne- Vaughan alone. The first of 
these, entitled “ Some Remarks on the Anatomy of the Osmundacese ” 
{Ann. of Rot., July 1911), dealt with the structure of the young plant, and 
the origin of the medullation, and of the foliar gaps in the individual life. 
The second was his last completed work, and it described a case of “mixed 
pith” found by Mrs G wynne- Vaughan in an anomalous specimen of 
Osmunda regalis. The structure seen in it was held to support the 
theory that the pith of the Osmundacese is phylogenetically stelar, and 
not cortical. 
The two series of Memoirs above mentioned embody the most effective 
work of G wynne- Vaughan. But they by no means exhaust it. He 
originated a most ingenious theory of the stele of Equisetum (Ann. of Rot., 
1901) ; he discovered the axillary buds of Helminthostachys (Ann. of Rot., 
1902) ; he worked through the anatomy of Archangiopteris (Ann. of 
Rot., 1905). He also wrote on the curious lattice-work structure of 
certain Fern stems (Ann. of Rot., 1905), and on the minute structure of 
the tracheae of Ferns, (Ann. of Rot., 1908). In all of these his originality 
was patent, though restrained. He had also entered upon other work in 
co-operation with Dr Kidston. A Memoir on Tempskya was published in 
Russia ( Verh . d. russ. kaiserl. mineral. Gesellschaft, Bd. xlviii, 1911), and 
a new series “ On the Carboniferous Flora of Berwickshire ” had been 
opened with its “Part I, Stenomyelon” (Trans. Roy. Soc. Edin., 1912). 
But here the curtain falls prematurely upon a life of investigation full of 
promise, as in the past it had been remarkably full of achievement. Only 
forty-four years of age, we can hardly forecast what G wynne- Vaughan 
might yet have done if he had lived out the full span. 
But he was not merely a laboratory botanist. He had an acute sense 
of specific characters, and a good knowledge of the native flora in the 
field. His systematic analysis of difficult species of Algse was pertinacious 
and successful,. And equally in the determination of Ferns, his systematic 
powers were exercised upon the collection brought back by Professor 
Lang from a journey in the Eastern Tropics. He was also an extensive 
traveller himself. In 1897 he went up the Amazon and Purus Rivers some 
3500 miles, as botanist attached to a rubber-prospecting expedition : but 
his scientific proclivities were restricted by the jealous demands of the 
firm that employed him. In 1899 he joined the Skeat Expedition to 
the Malay Peninsula, and experienced the delights of forest life with 
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