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PROFESSOR J. STEPHENSON AND DR BAINI PRASHA D ON 
directed obliquely upwards, towards the free upper edge of the lamella, and being 
more or less separated from each other do not form a regular palisade (fig. 9). 
They seem to be intermediate between typical rodlets and typical cilia, and again 
have some resemblance to the ragged hair-like appendages of the cells lower down 
on the lamellae. 
General Remarks on the foregoing Genera. 
So far there can be no doubt that the calciferous glands are to be considered 
as foldings of the oesophageal wall, and their epithelium as a continuation of the 
oesophageal epithelium. The condition described in Eutyphoeus is the most com- 
plicated ; the genus is to be derived from Octochsetus (though not directly — the 
intermediate stages having been lost, or not yet discovered) ; and the calciferous 
glands of Octochsetus have therefore been withdrawn, in Eutyphoeus , within the 
oesophagus instead of projecting as considerable diverticula at the sides. Looked at 
from another point of view, the glands of Eutyphoeus show us the extreme term 
attainable by a series of transverse lamellae ; their internal borders have fused 
together, so forming the vertical partition which separates off the interlamellar 
spaces from the general cavity of the oesophagus, and the communication of the 
interlamellar spaces with the oesophagus takes place only by means of their slit-like 
upper ends. 
The Calciferous Glands in the Lumbricida; (Figs. 11-13). 
Historical. 
It is in the Lumbricidae that the glands have been most fully investigated by pre- 
vious workers. Lankester was the first to examine them in any detail (15), though 
Morren had given a rough figure without any accurate description. Lankester calls 
them oesophageal glands, and places them in segments xii and xiii (corresponding to xi 
and xii in our present nomenclature). The first pair, in the first of the two segments, 
are round and full, very vascular, and firmly attached to the wall of the oesophagus, 
but do not appear to have any communication with the interior ; their wall is thin, 
and they contain each a single hard crystalline mass, or numerous smaller bodies ; the 
crystalline substance effervesces on the addition of acid. “ I have frequently found 
the crystalline bodies passed into the oesophagus and lodged in the capacious crop.” 
The second and third pairs of glands are both placed in the next segment, and are a 
little smaller than the first pair ; their walls are much thicker, but no less vascular ; 
they contain a milky secretion. In section, there are seen an inner epithelial coat, a 
vascular region, and an outer more delicate membrane forming the sheath of the organ, 
on which the externally visible vessels extend ; these vessels are shown in the figure 
as running longitudinally, parallel and numerous on all three pairs of glands. 
Claparede (5) identified three pairs of lateral pouches, the first two pairs in 
segment xi, the third in segment xii. He describes the glands as consisting of many 
