UNITED STATES DEPARTMENT OF AGRICULTURE 
In Cooperation with the 
Montana Agricultural Experiment Station 
DEPARTMENT BULLETIN No. 1403 
Washington, D. C. 
May, 1926 
SEGREGATION AND CORRELATED INHERITANCE IN MARQUIS AND HARD 
FEDERATION CROSSES, WITH FACTORS FOR YIELD AND QUALITY OF 
SPRING WHEAT IN MONTANA 
By J. Allen Clark, Agronomist in Charge, and John R. Hooker, Scientific 
Aid, Western Wheat Investigations, Office of Cereal Crops and Diseases, Bureau 
of Plant Industry 1 
CONTENTS 
Page 
Larger yields of wheat with present quality 
desired 1 
Material and methods 2 
Parent material 2 
Methods used 4 
Segregation of characters in the F2 and Fs 
generations.. 6 
Dwarfness 6 
Awnedness 9 
Glume color 16 
Kernel color 17 
Heading period 19 
Ripening period 26 
Fruiting period 33 
Height 39 
Page 
Segregation of characters in the F2 and F3 
generations— Continued . 
Yield 44 
Crude-protein content 49 
Combined characters 55 
Correlation of characters in the F2 and F3 
generations 57 
Heading period 58 
Ripening period 60 
Fruiting period 62 
Height 63 
Yield 64 
Crude-protein content 66 
Summary 67 
Literature cited 69 
LARGER YIELDS OF WHEAT WITH PRESENT QUALITY DESIRED 
The profitable production of hard red spring wheat on the dry- 
lands 01 Montana is dependent in part upon improvement in yield 
of the varieties grown. At present Marquis is the leading commercial 
spring wheat of the State. Under favorable conditions it is a high- 
yielding variety, but under conditions of deficient moisture Marquis 
is not adapted sufficiently to produce profitable crops. In unfavor- 
able seasons and in large sections having limited rainfall, therefore, 
spring-wheat production often is unprofitable in Montana, partly 
because of dependence on this or any other available variety. 
The quality of Marquis is satisfactory. The low-yielding Marquis 
grown on the Montana dry lands is especially high in quality and 
much desired by the grain trade. The problem is therefore to 
1 The Montana Agricultural Experiment Station cooperated in these investigations. Acknowledgment 
is gladly given here for field assistance rendered by M. A. Bell at Bozeman (1923) and Havre (1924), R. W. 
May at Moccasin, and J. L. Carter at Bozeman (1924). The writers also wish to express appreciation to 
Sewall Wright, of the Bureau of Animal Industry, United States Department of Agriculture, for advice 
given in the genetic and statistical analysis of the data. 
79182— 26t 1 
