FACTORS FOB YIELD AND QUALITY OF SPUING WHEAT 
55 
principally in the lower percentages. The data as a whole indicate 
that crude-protein content is inherited in intermediate degrees 
between the parents, through multiple factors. There is little or 
no evidence to show that transgressive segregation has occurred for 
higher protein strains than Marquis. Because of the fact that high 
crude-protein F 3 strains were found less variable than low crude- 
Erotein strains and the Marquis parent, it is hoped that selection of 
igh crude-protein F 3 plants may result in more uniform F 4 strains 
than Marquis and approach or equal it in average crude-protein 
content. 
COMBINED CHARACTERS 
No linkage was found between the four morphological characters — 
dwarfness, awnedness, and glume and kernel color — when the data 
on segregation were studied together. 
The four quantitative characters — heading period, ripening period, 
fruiting period, and height — which were studied to determine their 
effect on yield and crude-protein content, are more or less associated, 
and the nature and amount are determined by the correlation studies 
which follow. 
Table 50. — Average yield per plantX crude-protein content in the F 3 generation of 
521 Marquis-Hard Federation wheat hybrids, in comparison with 61 parent checks, 
in percentages of the average of Marquis at Bozeman and Havre, Mont., in 1924 
Yield X crude protein in percentages of the 
Marquis checks 
230 per cent . 
220 per cent . 
210 per cent . 
200 per cent . 
190 per cent . 
180 per cent . 
170 per cent . 
160 per cent . 
150 per cent . 
140 per cent . 
130 per cent . 
120 per cent . 
110 per cent. 
100 per cent . 
90 per cent . . 
80 per cent . . 
70 per cent . . 
60 per cent.. 
50 per cent . . 
40 per cent . . 
30 per cent . . 
Total 
Average per cent 
Bozeman 
F 3 
hybrids 
Hard 
Federa- 
tion 
Marquis 
1 i 
2 
3 
8 
21 
14 
32 
34 
42 
29 
18 
10 
7 
2 
1 
1 
1 
2 
1 
1 
2 
3 
4 
1 
1 
2 
2 
4 
1 
224 
36.4 
13 
81.9 
13 
100.0 
Havre 
hybrids I Mar ™ 
Hard 
Federa- 
tion 
9 
1 
9 
2 
12 
1 
23 
2 
1 
29 
2 
2 
49 
1 
2 
58 
1 
3 
48 
3 
4 
28 
1 
2 
8 
3 
1 
1 
297 
106.1 
17 
100.0 
18 
106.4 
1 Actual. 
Yield and crude-protein content so far have been considered inde- 
pendently of each other. From an agronomic standpoint high yield 
is the end result of an improvement program. The senior writer (5) 
has held that high yield "is not the only consideration; the quality 
of the product is of equal importance to that of yield." On this 
hypothesis the F 3 hybrids and parents have been studied on the 
basis of the product of yield per plant and crude-protein content. 
