INHERITANCE OF WAXY ENDOSPERM IN MAIZE. ae 
G and R are the presence of color factors and er their absence, both 
C and R being necessary to produce color. Were this true, wherever 
cR fertilized CR a colored seed would result, and this seed when 
planted would make the gametes OR and ch, which would give a 
ratio of 3 colored to 1 white, when self-pollinated. 
Wherever c? fertilized Or, a colored seed would result, wnich when 
planted would make four kinds of gametes as follows: Ch, Cr, ch, 
and cr. Self-pollinating such a plant would result in an ear with 9 
colored to 7 white seeds. 
In the second cross (Dh 234) it is assumed that the Chinese parent 
had the gametic composition of Cr Cr, while the Algeria parent had 
the gametic combination of CR CR. Wherever Cr fertilized Ch, a 
colored seed would result, which when planted would make the 
gametes CR and Cr. Such a plant when self-pollinated would give 
8 colored seeds to 1 white. As all the seeds of the hybrid Dh 284 
have the combination CC’ Rr, nothing but 3 to 1 ratios could result 
from self-pollinating any of the plants of this hybrid. 
When the combination CR Cr is pollinated with the other hybrid 
combination of CR cR, an all-colored ear would be the result, but when 
the combination of Dh 234 CR Cr is pollinated by the other combi- . 
nation of Dh 237, CR, Cr, cl, or er, an ear having 3 colored seeds 
to 1 white would be the result. This fulfills the observed conditions. 
Whenever an Algeria plant with the gametic composition CR Cr is 
self-pollinated, there should result an ear with 3 colored seeds to 1 
white. Nothing but all-colored ears have been obtained as a result 
of self-pollinating Algeria plants, but there have not been more 
than 10 or 12 self-pollinated Algeria plants, so that the failure to 
obtain an ear with 3 colored seeds to 1 white calls for no comment. 
Two other crosses described below do afford evidence, however, that 
Chinese plants of the gametic composition c? cP and Algeria plants 
with the composition CR Cr do occur. On the other hand, whenever 
a Chinese plant producing Cr gametes was pollinated with Chinese 
producing cl gametes an all-colored ear should result. We have 
never obtained any colored seed in crosses between two plants of 
white Chinese, although a large number of such crosses have been 
made, a fact which makes necessary the further assumption that 
plants with the gametic formula ascribed to one of the Chinese par- 
ent plants must be rare. 
An F, plant of the hybrid Dh 234 was fertilized with pollen from 
a white waxy Chinese plant; the resulting ear was all colored. A 
second ear on the Chinese plant was Ore pained and was all 
white, while a second ear on the hybrid Dh 284 was self-pollinated, 
producing 26 per cent white. These results accord with the idea 
89356°—19—_3 
