UNITED STATES DEPARTMENT OF AGRICULTURE 
DEPARTMENT BULLETIN No. 1429 
Washington, D. C. 
,1927 
THE PARASITES OF POPILLIA JAPONICA IN 
JAPAN AND CHOSEN (KOREA) AND THEIR 
INTRODUCTION INTO THE UNITED STATES 
By 
CURTIS P. CLAUSEN, -Entomologist 
J. L KING, Entomologist 
Japanese Beetle Investigations, Bureau of Entomology 
CHO TERANISHI, Assistant 
CONTENTS 
Page 
The General Problem . . 1 
The Field of Inrestigations 2 
The Parasites and Their Biology 3 
Centeter cinerea Aldrich 4 
Ochromeigenia ormioides Townsend .... 17 
Eutrixopsis javana Townsend ....... 19 
Prosena siberita Fabricius 20 
Dexia TenUalis Aldrich 26 
Campsomeris annulata Fabricius 31 
Tiphia popilliavora Rohwer 33 
Tiphia vernalis Rohwer 40 
Tiphia koreana Rohwer 42 
Craspedonotus tibialis Schaum ...... 43 
Page 
Life History of PopiUia japonica in Japan. . . 44 
Food Plants of Popillia in Japan. 47 
Climatic Conditions in the Main Fields of In- 
vestigation 48 
Agriculture and Natural Lands of Japan and 
Chosen in Relation to Popillia 51 
The Species of Popillia in Chosen . .... 53 
Popillia atrocoerulea Bates 53 
Popillia castanoptera Hope 53 
Popillia mutans Newm 54 
Summary 54 
WASHINGTON 
GOVERNMENT PRINTING OFFICE 
1927 
