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UNITED STATES DEPARTMENT OF AGRICULTURE 
DEPARTMENT BULLETIN No. 1468 
Washington, D. C. T March, 1927 
VARIABILITY IN THE LINKAGE OF TWO 
SEED CHARACTERS OF MAIZE 
G. N. COLLINS, Senior Botanist in Charge, and 
J. H. KEMPTON, Associate Botanist, Biophysical Laboratory 
Bureau of Plant Industry 
CONTENTS 
Page 
Introduction 1 
Statistical Treatment 4 
Variability of Crossover Rate 8 
Rate of Crossing Orer between the C-Wx Genes 24 
Correlations j, 56 
Discussion 60 
Summary 62 
Literature Cited * 64 
UNITED STATES 
GOVERNMENT PRINTING OFFICE 
WASHINGTON 
1927 
