News, Notes, and Reviews 
175 
in disintegrating cells they are very persistent elements of the 
cytoplasm. 
The nucleus of the carpogenic cell is succeeded by two nuclei, 
after which the nucleus of the trichophoric cell migrates down 
close to the carpogonium and undergoes a homoeotypic division. 
About the same time the septum between the two cells disappears, 
leaving a long four-nucleated cell. A restored trichophoric cell 
is then cut off carrying one of the trichophoric daughter nuclei; 
an inferior supporting cell with one of the two nuclei which suc- 
ceeded the carpogenic nucleus is cut off below. Four nuclei now 
appear in place of the two remaining centrally-placed nuclei, and 
a superior binucleate supporting cell is formed above, leaving a 
binucleated ascogonium below. The ascogonium may become an 
ascogenic cell directly, or a second inferior supporting cell may be 
cut off from it, and a second ascogenic cell may be formed, each 
cell being binucleate. At this important point the exact line of 
descent of the nuclei to be found in this series of binucleate cells 
was not determined. Whether the nuclei in the ascogenic cells 
are sister nuclei, a few divisions removed from the nucleus of 
the carpogonium, or whether they are non-sister nuclei, the great- 
granddaughters of the trichophoric and carpogenic cells respec- 
tively, is not stated. The author would seem to infer that the 
latter is the case by the statement that the nucleus of the tricho- 
phoric cell “ joins forces with that of the carpogonium.” The 
nuclei in the ascogenic cell divide conjugately, non-sister nuclei 
passing up into the ascus which buds off directly from the asco- 
genic cell. Their fusion follows. Four chromosomes appear at 
the conjugate divisions in the ascogenic cells and the same number 
is to be found at each division in the ascus ; reduction occurs in 
synapsis of the first division. A well-marked centrosome is found 
upon the nuclear membrane. The behavior of the centrosome in 
nuclear division, and its connection with the chromatin threads 
seems to be very similar to what Harper describes for the Erysi- 
phaceae. Faull still maintains, however, that the astral rays take 
no part as such in the delimitation of the spores, but agrees that 
the centrosome is the center of activity to the extent that the 
spores’ are delimited by the differentiation of a limiting layer 
