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POPULAK SCIENCE REVIEW. 
to leave some of its pollen adhering to the viscid organ. The 
stigma does not, as a rule, come so close to the lower lip as to 
be struck by the bee’s head ; this passes underneath it, and 
therefore it would be useless for the bee to carry pollen on its 
head. The sterility of the lower anther cells thus receives its 
explanation. 
I think I have now shown that every portion of this flower, 
even to such trifling peculiarities as might have been supposed 
to be without import, is really so modified as to facilitate ferti- 
lisation by the aid of bees, and to ensure intercrossing. I will 
briefly recapitulate the chief points. 
1. The Corolla, (a.) The lower lip affords a tempting landing- 
place on its middle lobe, while its side lobes regulate the direc- 
tion in which the bee settles, and compel it to approach the 
tube in a direct manner. {^.) The deep tube of the corolla 
compels the bee to dive into it for a certain depth, to get at the 
nectary, and so ensures the necessary pressure on the projecting 
anther cells. (7.) The secretion of nectar at the glandular base 
of the corolla attracts the bee by its taste and odour. (8.) The 
upper lip forms a close hood, so as to shelter the fertile anther 
cells, and prevent waste of pollen. The same lip, owing to its 
being curved and closed behind, compels the style to grow in 
such a form as eventually to bring its stigma into contact with 
the pollen-smeared back of the bee, (s.) The corolla bulges 
behind in such a way as to allow of free motion to the barren 
anther cells. 
2. The Stamens, (a.) The filaments are short and firm, so as 
to afford a secure point on which the connective may revolve. 
This security is increased in one salvia, if not in more, by adhe- 
rence of the filament to the upper lip of the corolla. {$.) The 
straddling position of the filaments leaves a free entrance for 
the bee. (7.) The filament is articulated with the connective 
so as to allow of a larger range of motion. (8.) The arm of 
the levers on which the bee directly acts is the shorter, so that 
the fertile cells are made to move considerably by a compara- 
tively slight motion of the bee itself, (e.) The lower anther 
cells are barren or nearly so, as pollen on the bee’s head would 
be wasted. On the other hand, the upper cells are very fertile, 
and their dehiscence is on the side which is brought into con- 
tact with the bee’s body, and which is turned away from the 
stigma, (f.) The lower cells project so that the bee must of 
necessity strike them. 
.3. The Style, (a.) The stigma ripens after the anthers. ( 0 .) 
The stigma is so placed {is to be protected from the pollen of 
the same plant. (7.) The style grows in a curve, so as even- 
tually to come in contact with the back of the bee. 
I have described this variety of salvia at such length that it 
