800 Proceedings of Royal Society of Edinburgh. [sess. 
contains elongated tubular glands embedded in a small amount of 
connective tissue, in which is a system of delicate blood-vessels and 
lymphatics. Between the tubular glands in the deeper layer the 
amount of connective tissue is very slight indeed, but near the 
surface epithelium, where the gland ducts proper are situated, it 
becomes more abundant. The glands are simple unbranched tubes 
arising each from a duct of its own, and running almost straight 
from the epithelial surface to the submucosa, where it ends in a 
short hooked extremity. The secreting portion of the walls of these 
glands is formed of a double layer of cells (Plate II. fig. 2, and 
Plate III. figs. 1 and 2). Internally, a layer of cells (viz., the central 
cells of the gland) are for some distance down columnar in form, but 
they become smaller in size and distinctly cubical as we come to the 
deeper part of the gland, the amount of protoplasm in each cell 
gradually becoming smaller. Outside this is a second layer of cells 
(the parietal cells of the gland), completely investing the tubule, 
and not occurring merely at intervals as in the cardiac glands of the 
stomach of the dog and human subject. The cells are large nucleated 
protoplasts, irregular in shape, slightly flattened, pyramidal, &c., each 
lodged in a distinct cavity formed by a framework of delicate con- 
nective tissue on which small flattened nuclei are seen (Plate IY. 
fig. 1). These cells are largest in the deepest part of the mucosa; 
near the free surface they become smaller in size, more flattened, and 
eventually they disappear. 
This second compartment corresponds with the first compartment 
of the Ziphioid whales as described by Turner* in Mesoplodon 
bidens and in Hyperoodon rostratus , and is in the Narwhal 
somewhat larger than the first compartment without the diver- 
ticulum, though scarcely so much as one would gather from Meckel’s 
description. Murie,f on the other hand, says that in Globio- 
cephalus the first gastric cavity is by far the largest. 
With it commences the true secreting portion of the stomach, 
and it may be compared to the cardiac end of the dog’s stomach. 
The point of special interest is that the peripheral cells are 
specially numerous, forming a layer completely investing the 
central or axial columnar epithelial cells. The reticulum, in 
which the large parietal cells are embedded, corresponds apparently 
* (5) Page 247. t (6) Page 247. 
