802 Proceedings of Royal Society of Edinburgh. [sess. 
partment were lined by a double set of cells — the “ central ” and 
“ parietal.” Of these in this cavity, in most cases ordinary columnar 
epithelial secreting cells, resting on a basement membrane of flat- 
tened nucleated cells, and corresponding to the “central” cells, 
alone are present (Plate I. fig. 2); but in a few glands, at what 
may be called the cardiac extremity, a few of the large parietal 
cells are found in the deeper part of the tube (Plate IY. fig. 2). 
The framework of connective tissue supporting the glands is 
much more abundant and stronger than in the previous compart- 
ment. 
It is generally agreed that in Delphinidse this portion of the 
stomach is much narrower and more intestine-like than any of the 
other compartments. Meckel, speaking of this portion of the 
stomach of the Narwhal, describes it as much narrower and longer, 
commencing with a small caecum, which lies backwards and to the 
left, first taking a turn forwards, then quickly backward, then again 
forward and somewhat to the right, in order to join by a very 
narrow sphincter orifice the duodenum. (This can scarcely corre- 
spond to the cavity above described in anything but shape.) In 
the case of Globiocephalus, Murie * describes it as a communicat- 
ing passage, which “ leaves the second stomach on its right inferior 
wall, an inch below the wide aperture which connects the first and 
second, and it enters the third stomach above and behind. There 
is no true sphincter at either end.” 
Turner,! referring to Murie’s description, says that “in Globio- 
cephalus melas, both Dr Jackson and I have described five com- 
partments, though Dr Murie regards the compartment which I have 
numbered three not as a true digestive sac, but only as a communi- 
cating canal. With this interpretation, however, I am unable to 
agree, and still adhere to my opinion that it is a true, though 
small, gastric compartment.” Morrison Watson and Young 
give their reasons for dissenting from Murie’s view briefly, as 
follows : — 
They point out that in many animals, e.g., birds, certain limited 
areas of the stomach are set apart for special glandular secretion. 
* Trans. Zool. Soc. Lond., vol. viii. part 4, p. 257. 
t “Anatomy of Sowerby’s Whale,” Jour. Anat. and Phys.. October 1885, 
p. 154. 
