698 Proceedings of the Royal Society of Edinburgh. [Sess. 
and of studying in fresh specimens the hand with the bones in situ , and 
not artificially articulated. I propose therefore to continue my account of 
the morphological constitution of the manus in the toothed whales.* 
Hyperoodon rostratus. 
A young male 204 feet long was stranded near Dunbar in November 
1885. I secured the specimen and mounted the skeleton in the Anatomical 
Museum of the University. As the vertebral plates and the epiphyses of 
the long bones were not fused with their diaphyses the animal had not 
reached maturity. The carpal end of both radius and ulna had a well- 
marked epiphysial cartilage partially ossified. The carpalia were flattened 
on the surfaces and consisted of osseous nodules in masses of cartilage. 
The proccirpus or proximal row contained radiale, intermedium and 
ulnare. The radiale show T ed an osseous nodule 31 mm. in transverse 
diameter surrounded by cartilage which articulated with radius, intermedium 
and ulnare. In the intermedium the osseous nodule was 41 mm. wide, and 
with its surrounding cartilage articulated with radius, ulna, radiale, ulnare 
and carpalia 2 , 3 , 4 . The osseous nodule in the ulnare was 32 mm. wide, and 
with its cartilage articulated with ulna, intermedium, carpalia 4 , 5 , and with 
the unossified pisiform cartilage, which formed the inner border of the carpus 
and was connected with the cartilage of the ulnare, carpale 5 and metacarpal v . 
The distal row, mesocarpus, consisted of five distinct elements, three of 
which were partially ossified and two were unossified. Carpale j had an 
osseous nodule 17 mm. wide, and articulated with radiale, C 2 and M . 
Carpale 2 was unossified and articulated with radiale, intermedium, C 1? 
C 3 and M n . Carpale 3 had an osseous nodule 24 mm. wide, and 
articulated with intermedium, C 2 , C 4 and Mm. Carpale 4 had an osseous 
nodule 16 mm. wide, and articulated with intermedium, ulnare, C 3 , C 5 and 
Miy. Carpale 5 was unossified and articulated with ulnare, pisiform cartilage, 
C 4 and My. The carpal elements in this Hyperoodon, though imperfectly 
ossified, corresponded in number and position with the bony carpus of the 
adult female previously described and a figure f of which is now reproduced 
on the next page ; they showed that ossification did not proceed uniformly in 
the bones, for a needle passed through C 2 and C 5 failed to detect bone in 
the substance of their cartilage, whilst in C l5 C 3 and C 4 the osseous nodule 
was visible on the surface. No os centrale was present in either carpus. 
The hand was pentadactylous, the metacarpal of each digit articulated 
* In the Proc. Roy. Soc. Edin ., 1891-92, vol. xix. p. 70, I have discussed the 
morphology of the manus in the whalebone whale, Balcenoptera rostrata. 
t Journ. of Anat. and Phys., October 1885, vol. xx. p. 184. 
